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The main shallow water marine and coastal habitats: (A) Cuvumbi Island reef flat, showing Acroporacolonies during lowest low tide; (B) wide abraded flat bordering the Lac Badana outlet, emerged at low tide. The rugged surface is covered by a thin sandy sheet and scattered seagrass; (C) mangal flat at low tide near Yamani (Lac Badana). The roots are densely encrusted by ostreids (Saccostrea
cuccullata); (D) Fuma Island sandy tail at low tide showing a densely bioturbated
muddy bottom; (E) ebb-oriented ripples on the beach ridge at Lac Anole outlet