Program Abstracts 4th WDA Symposium PDF

Abstracts Book 4th WDA International Symposium of Odonatology, Pontevedra 26-30 July 2005
Program and Abstracts
Edited by Adolfo Cordero Rivera, University of Vigo
Organizing commitee: Adolfo Cordero Rivera (organizer), Universidade de Vigo Csar Ayres Fernndez Josefina Garrido Gonzlez Ins Gonzlez de Castro Olalla Lorenzo Carballa Isabel Pardo Gamundi Rosa Ana Snchez Guilln Guillermo Velo Antn Postal address: Laboratorio de Ecoloxa, EUET Forestal, Universidade de Vigo, Campus Universitario, 36005 Pontevedra, Spain. Scientific Commitee Commitee member Adolfo Cordero Rivera Alejandro Crdoba Aguilar Ola M. Fincke Josefina Garrido Gonzlez Manuel Graa Isabel Pardo Gamundi Michael J. Samways David J. Thompson University Vigo, Spain UNAM, Mxico Oklahoma, USA Vigo, Spain Coimbra, Portugal Vigo, Spain Matieland, South Africa Liverpool, UK
Tuesday 26 July
From 10:00 10:30 11:00 11:30 12:00 12:30 13:00 15:00 15:30 16:00 16:30 17:30 18:00 18:30 19:00 To 10:30 11:00 11:30 12:00 12:30 13:00 15:00 15:30 16:00 16:30 17:30 18:00 18:30 19:00 19:15 Forests and dragonflies Forests and dragonflies Forests and dragonflies Informal Forests and dragonflies Forests and dragonflies Forests and dragonflies Taylor, Ph. Paulson, D. Fincke, O.M. Forests and dragonflies Forests and dragonflies Samways, M. Sahln, G. Topic Opening ceremony Forests and dragonflies Forests and dragonflies Authors Title of communication
Corbet, P.S. Orr, A.G.
Forests as habitats for dragonflies (Odonata) Odonata in Bornean tropical rain forest formations: diversity, endemicity and implications for conservation management Coffee break Threat levels to odonate assemblages from invasive alien tree canopies Specialists vs. generalists among dragonflies - the importance of forest environments to form diverse species pools Lunch Movement behaviors of odonates in heterogeneous forest landscapes The importance of forests to neotropical dragonflies Habitat use by pseudostigmatid damselflies: their future in fragmented forests Coffe break (poster installation)
Claustnitzer, V. & What matters to tropical forest dragonflies? African impressions Dijstra, K.-D. Historical and recent population genetics: Any news to tell us about Hadrys, H. the impact of fragmentation on forest odonates? Thompson, D.J. & The structure of the Coenagrion mercuriale populations in the New Watts, Ph.C. Forest, southern England Reflections on the 2003 International Odonatology Symposium, Hawking, J.H. Beechworth, Australia
Wednesday 27 July
From 9:30 10:00 10:15 10:30 10:45 11:00 11:45 12:00 12:15 12:30 12:45 15:00 15:15 15:30 15:45 16:00 16:15 16:30 17:30 17:45 18:00 18:15 18:30 18:45 19:00 To 10:00 10:15 10:30 10:45 11:00 11:45 12:00 12:15 12:30 12:45 15:00 15:15 15:30 15:45 16:00 16:15 16:30 17:30 17:45 18:00 18:15 18:30 18:45 19:00 19:15 Ecology and conservation Ecology and conservation Ecology and conservation Ecology and conservation Ecology and conservation Ecology and conservation Informal Sexual selection Sexual selection Sexual selection Sexual selection Topic Forests and dragonflies Behaviour Behaviour Behaviour Behaviour Behaviour Behaviour Behaviour Behaviour Authors Watanabe, M. Fincke, O.M. Title of communication Mate location and competition for mates in relation to sunflecks of forest floors Lack of innate recognition of species or morph identity in Enallagma damselflies
Van Gossum, H.; Beirinckx, K.; Large-scale variation in female morph frequencies of the polychromatic damselfly Nehalennia irene Forbes, M. & Sherratt, T. Hilfert-Rppell, D. Documenting odonate behaviour by drawing from films Ovipositor and egg-laying behaviour of Odonata: phylogenetic Mathuskina, N. implications Coffee break and poster session Rppel, G. Flashes in flight communication between odonate males Snchez-Guilln, Pre- and postcopulatory mechanisms of reproductive isolation R.A. & Cordero between Ischnura graellsii and I. elegans Rivera, A. Does ovaries composition vary between species with different mateSchenk, K. guarding intensities? Olberg, R. & Dragonfly prey capture: Vision, decision, and flight Worthington, A. Lunch Copulatory behaviour in hybrid matings between Calopteryx Cordero Rivera, A. haemorroidalis and C. splendens Crdoba-Aguilar, A.Sperm ejection as a cryptic female choice mechanism in odonates Serrano-Meneses, Survival and mating success of American Rubyspots in relation to M. A.; Szkely, T. & body size (Odonata: Calopterygidae) Crdoba-Aguilar, A. Szllassy, N.; Differences in survival of mated and unmated males of Libellula Szab, Z.D. & fulva: a four year study Nagy, H.B. Female reproductive behaviours of different Chlorocyphid species in Gnther, A. the Oriental-Australian region The global dragonfly assessment
Sexual selection Monitoring and Spector, S. climate change
Coffee break, poster session and IUCN Odonata Specialist Group meeting Why are endemic Odonates are endangered in the oceanic islands Karube, H. of Ogasawara? Riservato, E. & Dragonflies of riverine habitats: assessment as indicators of Bogliani, G. biodiversity and environmental integrity Kadoya, T.; Suda, Spatial heterogeneity of the dragonfly assemblages in the S.; Washitani, I. & landscape scale: assessments using newly created small ponds as Tsubaki, T. traps in the catchment area of lake Kasumigaura Lorenzo Carballa, M.O. & Cordero Fecundity and fertility of parthenogenetic Ischnura hastata Rivera, A. Susa, K. & Habitat selection and egg production in Sympetrum infuscatum Watanabe, M. females living in a forest-paddy field complex Martens, A. & Ecology of Odonata inhabiting permanent Namibian desert springs Suhling, F. Suhling, F. & Desperately seeking the man in the upside-down bucket - some Sahln, G. recollections from the WDA symposium in Beechworth, Australia
Thursday 28 July Mid-symposium trip to Corrubedo Natural Park. Buses will depart from the Forestry School at 9:00 and return at about 19:30. Itinerary: visit to the Hydrobiology Field Station of the University of Santiago, and Corrubedo Natural Park, visiting the big dune (16 m of maximum elevation; more than one km in length), and a number of small ponds and channels. There are two coastal lagoons in the park (Carregal and Vixn). The first is permanently connected to the sea, and has very few dragonflies. The lagoon of Vixn, with less that 2 m of depth, has a rich fauna of aquatic insects. A pre-roman settlement ("Castro of Baroa") is in the vicinities, and could be visited in the afternoon. It will be possible to go to the beach and have a bath, but take into account that water temperature might be around 14C.
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Friday 29 July
From 9:30 11:00 11:45 12:00 12:15 12:30 12:45 15:00 15:15 15:30 15:45 16:00 To 11:00 11:45 12:00 12:15 12:30 12:45 15:00 15:15 15:30 15:45 16:00 16:15 Ecology Ecology Life-history Life-history Topic Authors May, M.L. (coordinator) Title of communication Plenary seminar
Coffee break and poster session Watanabe, Y. Artificial parthenogenesis in Aeshna nigroflava Martin Weihrauch, F.; Olias, Distribution and overlap of ranges of Lestes parvidens and M.; Bedjani, M.; Marinov, M. & Lestes viridis in southeastern Europe (Odonata: Lestidae) alamun, A. Odonata larvae and drought in Australia: Ecological Hawking, J.H. development for life in an unpredictable climate The influence of environment and phylogeny on the Suhling, F. & Sahln, determination of morphological, behavioural and life history G. traits in dragonfly larvae Lunch Quaternary environmental change along the Western Escarpment of Africa and the distribution of Namibian Odonata The occurrence of Cordulegaster sp. in Czech Republic result of influence of habitat ecological factors in different biogeographical regions? Notes on Crimean Odonata (Crimea, Ukraine) Biogeography and habitat affinity of the odonata fauna of SE Madagascar Population differences in sexual selection intensity and immune response in two contrasting forest environments in the damselfly Hetaerina americana (Zygoptera: Calopterygidae) Southern dragonflies expanding in Wallonia (South Belgium) : a consequence of global warming? Coffee break and poster session
Faunistics and Marais, E. distribution Faunistics and Holua, O. distribution Faunistics and Khrokalo, L. & distribution Prokopov, G. Faunistics and Schtte, K. distribution Contreras-Garduo, J. & CrdobaAguilar, A. Goffart, Ph., Fichefet, V., De Monitoring and Schaetzen R., climate change Baugnee J-Y., Lebrun, Ph. & Dufrene, M. Forests and dragonflies Bouwman, J.; Monitoring and Groenendijk, D. & climate change Plate, C. Monitoring and Conze, K.-J. climate change Monitoring and De Knijf, G. & climate change Anselin, A. Monitoring and Kalkman, V. climate change
16:15
16:30
16:30 17:30 17:45 18:00 18:15 18:30 20:45
17:30 17:45 18:00 18:15 18:30 19:45 ??
The Dutch Dragonfly Monitoring Scheme: results and trends Dragonfly monitoring in northrhine Westfalia, Germany When south goes north: mediterranean Odonata conquer Flanders (North Belgium) Towards an atlas of European odonates WDA Biennial General meeting Symposium dinner
Saturday 30 July
From 9:30 10:00 10:15 10:30 10:45 11:00 11:45 12:00 12:15 12:30 12:45 15:00 15:15 To 10:00 10:15 10:30 10:45 11:00 11:45 12:00 12:15 12:30 12:45 15:00 15:15 15:30 Topic Forests and dragonflies Ecological genetics Title of communication Mapping potential habitats using environmental surrogate Tsubaki, Y. measures: Importance of forests for dragonflies in Japan Giere, S. & Hadrys, Genetic consequences of habitat specialisation and cryptic H. speciation in the genus Trithemis The effects of Climatic Changes for the distribution of Monitoring and Ott, J. dragonflies in Europe and their possible effects on the climate change biocoenosis of the waters Monitoring and Ubukata, H. & Optimization of environmental monitoring schedule using adult climate change Sakoda, T. dragonflies Climate impacts on a North American dragonfly: evolutionary vs. Monitoring and Matthews, J.H. climate change ecological responses Coffee break Flying Colours: Five years of research on Odonata in tropical Systematics and Dijkstra, K.-D. eastern Africa Morphology Systematics and Kjer, K.M.; Carle, A preliminary phylogenetic hypothesis of Odonata, based on Morphology F.L. & May, M.L. multiple molecular and morphological data sets Systematics and Leipelt, K.G. Ecomorphology of legs in larval and adult Anisoptera Morphology Critical and consequent taxonomy in Odonata: the European Systematics and Dijkstra, K.-D. perspective Morphology Lunch Aquatic egg-parasitoids (Hymenoptera) of dragonflies and other Miscellanea Fursov, V. arthropods: unique life and flight under water On some paintings of Odonata from the late Middle Ages (14th Miscellanea Carvalho, A.L. and 15th centuries) Sathe,T.V.; Basarkar, C.D.; Impact of dragonflies on population suppression of paddy pests Mundale, M.; Miscellanea in agroecosystem of Kolhapur district, India Bhosale, Y.A. & Margaj, G.S. Karube, H.; Katatani, On the genus Dubitogomphus Fraser, 1940, their true status Miscellanea N. & Kitagawa, K. and characters Manu Thomas, Comparative studies on the genital and sub-genital abdominal Morphology Gunasekaran & segments of five species of dragonflies (Anisoptera: Odonata) Mohan Daniel Allochthonous organic matter as a food resource for aquatic Closing talk Graa, M. invertebrates in forested streams Closing ceremony Authors
15:30
15:45
15:45 16:00 16:15 16:45
16:00 16:15 16:45 17:15
Sunday 31 July to Thursday 4 August, Postsymposium tour
List of poster presentations

Title of poster Preliminary observations of reproductive behavior in Arigomphus villosipes (Anisoptera: Gomphidae) 2 Behaviour Nagy, H.B.; Lszl, Z.; Szllassy, Site fidelity, mating success and reproductive strategies in males N.; Szkely, A. & Dvai, G. of Libellula fulva (Odonata: Libellulidae) 3 Behaviour Sathe, T.V.; Bhoje, P.M.; Kavane, Mating behaviour in damselfly Ischnura senegalensis (Rambur) P.; Shinde, K.; Yadav, R.P. & Pandharbale, A.R. 4 Behaviour Schenk, K. Egg distribution, mate-guarding intensity and offspring conditions in dragonflies 5 Behaviour Contreras-Garduo, J.; Buzatto, Fat reserves and wing spot size, but not wing spot and body colour B.; Najera, K. & Crdoba-Aguilar, intensity, are related to male success during territorial contests in A. Hetaerina americana 6 Conservation Mihokovic, N. & Slavikovski, A. The effect of thermal pollution on dragonfly populations 7 Conservation Rodrguez-Guntn, I.; PrezEvaluation of the odonata community in Galician rivers (NW Spain) Bilbao, A.; Gonzlez, A.; Alonso, that are affected by hydroelectric power stations A. & Garrido, J. 8 Ecological Timm, J. & Hadrys, H. Comparative molecular genetic analysis of the population genetics structures and dynamics of two aeshnid species (Odonata: Aeshnidae) in Namibia 9 Ecological Van Gossum, H.; Snchez Hybridisation and inheritance of female-limited colour genetics Guilln, R.A. & Cordero Rivera, A. polymorphism in two ischnurid damselfly species (Odonata) 10 Ecological Wargel, A.; Giere, S. & Hadrys, H. Genetic consequences of a man-made bottleneck in Orthetrum genetics coerulescens: A microsatellite system to study fine scale population dynamics 11 Ecology Gonzlez de Castro, I.; van The effect of larval density on female polychromatism and body Gossum, H. & Cordero Rivera, A. size in Ischnura graellsii 12 Ecology May, M.L. Preliminary results of a multi-year study of phenology and development of Anax junius in Maryland, U.S.A. 13 Ecology Snchez Guilln, R.A. & Cordero Relative frequency of Ischnura elegans and I. graellsii (Odonata: Rivera, A. Coenagrionidae) in the Galician coast 14 Faunistics Alonso, A.; Garrido, J.; PrezPresent dragonflies in As Gndaras de Budio. Site of Bilbao, A.; Gnzalez, A. & Community Importance (Nature Network 2000) Rodrguez-Guntn, I. 15 Faunistics Ferreira, S. & Grosso-Silva, J.M. Present knowledge on the Odonata of Serra da Estrela Natural Park (Portugal) 16 Faunistics Ferreira, S.; Grosso-Silva, J.M.; Odonata of continental Portugal: mapping the knowledge and Soares-Vieira, P. & Sousa, P. identifying geographical gaps 17 Faunistics Oppel, S. Dragonflies and damselflies of a montane tropical rainforest in Papua New Guinea 18 Faunistics Sathe, T.V.; Shinde, K.; Bhoje, Biodiversity of dragonflies (Odonata) from Western ghats of P.M.; Thite, H.S. & Patil, R.G. Maharashtra, India 19 Faunistics Soares-Vieira, P.; Grosso-Silva, On the available data concerning the Odonata of Peneda-Gers J.M. & Ferreira, S. National Park (NW Portugal) 20 Faunistics Azpilicueta, M.; Rey Ra, C.; A preliminary a analysis of odonate species richness in Galiza Docampo Barrueco, F.; Rey (NW Spain) Muiz, X.C. & Cordero Rivera, A. 21 Life-history Cano-Villegas, F.J. & Ferreras- Contribution to knowledge of the biology of Onychogomphus Romero, M. costae Slys, 1885 (Odonata: Gomphidae) in southern Spain 22 Life-history Torralba Burial, A. & Ocharan, F.J.Pond water regime and competition as key factors in the presence and life history of two Lestes damselflies (Odonata: Lestidae) 1 Topic Behaviour Authors McMillan, V.E.
Topic 23 Monitoring and climate change 24 Monitoring and climate change 25 Monitoring and climate change 26 Morphology 27 Pollution
Authors Khrokalo, L. & Matushkina, N. Kurauchi, Y. & Ubukata, H. Termaat, T.; Ketelaar, R. & de Vries, H. Watanabe, Y.
Title of poster Expansion of Crocothemis erythraea in Ukraine Sensitivity and economy of monitoring the environment of a large lake using adult dragonflies Flight peak trends for dragonflies from the Netherlands
28 Pollution
Morphological characteristics of odonate eggs and early Instar larvae specific to taxa Ferreras-Romero, M.; CanoInterannual change measurement of the Odonata community Villegas, F.J. & Rubio-Soler, M.I. existing in a Mediterranean river which put up with the by-product of a heavy mining accident happened in April 1998 (river Guadiamar, Andalusia, southern Spain) G. Rodrguez-Liares; Garrido, J.; Relationships between the concentrations of heavy metals in Bendicho, C. & Lavilla, I. dragonfly larvae and Louro river sediments G. Rodrguez-Liares; Garrido, J.; Evaluation of three species of dragonfly larvae as biomonitors of Bendicho, C. & Lavilla, I. heavy metals Fernndez Martnez, M.A. Dragonfly folklore in Galicia, Northwest Iberian Peninsula A new genus of Coenagrionidae (Odonata, Zygoptera) from the Pantepui region of Venezuela, with descriptions of four new species WAPODOKEY A comparative study of haemocytes in male and female of the dragonfly Crocothemis servilia Drury
29 Pollution 30 Sociology
31 Systematics De Marmels, J. 32 Taxonomy 33 Physiology Mihokovic, N. Ashok, J.; Shinde, K. & Sathe, T.V.
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Index Oral contributions........................................................................................................................... 15 Forests as habitats for dragonflies (Odonata) ............................................................................. Odonata in Bornean tropical rain forest formations: diversity, endemicity and implications for conservation management................................................................................. Threat levels to odonate assemblages from invasive alien tree canopies................................ Specialists vs. generalists among dragonflies - the importance of forest environments to form diverse species pools .............................................................................................................
Corbet, Philip S....................................................................................................................................................15
Orr, Albert G. ......................................................................................................................................................15 Samways, Michael J. ............................................................................................................................................16
Sahln, Gran ......................................................................................................................................................17 Taylor, Philip D. ..................................................................................................................................................18 Paulson, Dennis...................................................................................................................................................18 Fincke, Ola M. .....................................................................................................................................................19 Clausnitzer, Viola & Dijkstra, Klaas-Douwe "KD" B....................................................................................20
Movement behaviors of odonates in heterogeneous forest landscapes..................................
The importance of forests to neotropical dragonflies ............................................................... Habitat use by pseudostigmatid damselflies: their future in fragmented forests ................... What matters to tropical forest dragonflies? African impressions...........................................
Historical and recent population genetics: Any news to tell us about the impact of fragmentation on afrotropical forest odonates?.......................................................................... The structure of the Coenagrion mercuriale populations in the New Forest, southern England............................................................................................................................................. Mate location and competition for mates in relation to sunflecks of forest floors ...............
Hadrys, Heike ......................................................................................................................................................20
Thompson, David J. & Watts, Phillip C...........................................................................................................21 Watanabe, Mamoru.............................................................................................................................................22 Fincke, Ola M. .....................................................................................................................................................23
Lack of innate recognition of species or morph identity in Enallagma damselflies................ Large-scale variation in female morph frequencies of the polychromatic damselfly Nehalennia irene..................................................................................................................................
Documenting odonate behaviour by drawing from films......................................................... Ovipositor and egg-laying behaviour of Odonata: phylogenetic implications .......................
Van Gossum, Hans, Beirinckx, Kirsten, Forbes, Mark and Sherratt, Thomas...........................................23 Hilfert-Rppell, Dagmar.....................................................................................................................................24 Matushkina, Natalia.............................................................................................................................................25 Rppell, Georg.....................................................................................................................................................26
Flashes in flight communication between odonate males .....................................................
Pre- and postmating mechanisms of reproductive isolation between Ischnura graellsii and I. elegans (Odonata: Coenagrionidae)............................................................................................. Does ovaries composition vary between species with different mate-guarding intensities?
Snchez-Guilln, Rosa Ana & Cordero Rivera, Adolfo.................................................................................26 Schenk, Kamilla ...................................................................................................................................................27 Olberg, Robert and Worthington, Andrea.......................................................................................................27 Cordero Rivera, Adolfo ......................................................................................................................................28 Crdoba-Aguilar, Alex........................................................................................................................................29
Dragonfly prey capture: Vision, decision, and flight.................................................................. Copulatory behaviour in hybrid matings between Calopteryx haemorroidalis and C. splendens .
Sperm ejection as a cryptic female choice mechanism in odonates......................................... Survival and mating success of American Rubyspots in relation to body size (Odonata: Calopterygidae) ................................................................................................................................ Differences in survival of mated and unmated males of Libellula fulva: a four year study ....
Serrano-Meneses, M. A., Szkely, T. and Crdoba-Aguilar, Alex ................................................................29
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Female reproductive behaviours of different Chlorocyphid species in the OrientalAustralian region.............................................................................................................................. The global dragonfly assessment...................................................................................................
Szllassy, Nomi, Szab, Zoltan D., Nagy, H. Beta......................................................................................30
Gnther, Andr ...................................................................................................................................................31 Spector, Sacha; Naskrecki, Piotr........................................................................................................................31 Karube, Haruki ....................................................................................................................................................32
Why are endemic Odonates endangered in oceanic islands Ogasawara?................................
Dragonflies of riverine habitats : assessment as indicators of biodiversity and environmental integrity...................................................................................................................
Spatial heterogeneity of the dragonfly assemblages in the landscape scale: assessments using newly created small ponds as traps in the catchment area of lake Kasumigaura.........
Riservato, Elisa & Bogliani, Giuseppe..............................................................................................................33
Kadoya, Taku; Suda, Shin-ichi, Washitani, Izumi & Tsubaki, Yoshitaka ....................................................33 Lorenzo Carballa, Olalla & Cordero Rivera, Adolfo......................................................................................34
Fecundity and fertility of parthenogenetic Ischnura hastata ........................................................ Habitat selection and Egg production in Sympetrum infuscatum females living in a forestpaddy field complex ........................................................................................................................
Susa, Koichi & Watanabe, Mamoru..................................................................................................................35 Martens, Andreas & Suhling, Frank..................................................................................................................36 Watanabe, Yoko ..................................................................................................................................................37
Ecology of Odonata inhabiting permanent Namibian desert springs.....................................
Artificial parthenogenesis in Aeshna nigroflava Martin................................................................. Distribution and overlap of ranges of Lestes parvidens and Lestes viridis in southeastern Europe (Odonata: Lestidae)...........................................................................................................
Odonata larvae and drought in Australia: Ecological development for life in an unpredictable climate ......................................................................................................................
Weihrauch, Florian, Olias, Marko, Bedjani, Matja, Marinov, Milen & alamun, Ali..............................37
Hawking, John H.................................................................................................................................................38
The influence of environment and phylogeny on the determination of morphological, behavioural and life history traits in dragonfly larvae ................................................................
Quaternary environmental change along the Western Escarpment of Africa and the distribution of Namibian Odonata ............................................................................................... The occurrence of Cordulegaster sp. in Czech Republic result of influence of habitat ecological factors in different biogeographical regions?............................................................
Suhling, Frank & Sahln, Gran .......................................................................................................................39
Marais, Eugene ....................................................................................................................................................40
Notes on Crimean Odonata (Crimea, Ukraine).......................................................................... Biogeography and habitat affinity of the odonata fauna of SE Madagascar ..........................
Holua, Otakar.....................................................................................................................................................41 Khrokalo, Lyudmyla, Prokopov, G. .................................................................................................................42 Schtte, Kai ..........................................................................................................................................................42
Population differences in sexual selection intensity and immune response in two contrasting forest environments in the damselfly Hetaerina americana (Zygoptera: Calopterygidae) ................................................................................................................................ Southern dragonflies expanding in Wallonia (South Belgium) : a consequence of global warming?...........................................................................................................................................
Contreras-Garduo, Jorge & Crdoba-Aguilar, Alex.....................................................................................43
The Dutch Dragonfly Monitoring Scheme: results and trends. ...............................................
Goffart, Ph.; Fichefet, V.; de Schaetzen R.; Baugne J-Y.; Lebrun, Ph. & Dufrne, M. ...........................44 Bouwman, Jaap, Groenendijk, Dick & Plate, Calijn.......................................................................................44 Conze, K.-J...........................................................................................................................................................45 De Knijf, Geert & Anselin, Anny .....................................................................................................................46
Dragonfly monitoring in northrhine Westfalia, Germany......................................................... When south goes north: mediterranean Odonata conquer Flanders (North Belgium) ........
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Towards an atlas of European odonates...................................................................................... Mapping potential habitats using environmental surrogate measures: Importance of forests for dragonflies in Japan .....................................................................................................
Kalkman, Vincent................................................................................................................................................47
Tsubaki, Yoshitaka ..............................................................................................................................................48
Genetic consequences of habitat specialisation and cryptic speciation in the genus Trithemis .............................................................................................................................................
The effects of Climatic Changes for the distribution of dragonflies in Europe and their possible effects on the biocoenosis of the waters ...................................................................... Optimization of environmental monitoring schedule using adult dragonflies....................... Climate impacts on a North American dragonfly: evolutionary vs. ecological responses....
Giere, Sandra & Hadrys, Heike .........................................................................................................................48
Ott, Jrgen............................................................................................................................................................49 Ubukata, Hidenori and Sakoda, Tetsuo ...........................................................................................................50 Matthews, John H. ..............................................................................................................................................50 Dijkstra, Klaas-Douwe "KD" B........................................................................................................................51
Flying Colours: Five years of research on Odonata in tropical eastern Africa....................... A preliminary phylogenetic hypothesis of Odonata, based on multiple molecular and morphological data sets ..................................................................................................................
Kjer, Karl M.; Carle, Frank L. & May, Michael L. ..........................................................................................52 Leipelt, Klaus Guido...........................................................................................................................................52 Dijkstra, Klaas-Douwe "KD" B........................................................................................................................53
Ecomorphology of legs in larval and adult Anisoptera ............................................................. Critical and consequent taxonomy in Odonata: the European perspective ........................... Aquatic egg-parasitoids (Hymenoptera) of dragonflies and other arthropods: unique life nd flight under water.......................................................................................................................
On some paintings of Odonata from the late Middle Ages (14th and 15th centuries) ........... Impact of dragonflies on population suppression of paddy pests in agroecosystem of Kolhapur district, India. .................................................................................................................
Fursov, Viktor......................................................................................................................................................54 Carvalho, Alcimar do Lago ................................................................................................................................55
On the genus Dubitogomphus Fraser, 1940, their true status and characters ............................
Sathe, T.V.; Mundale, Mandar; Bhosale, Y.A. & Margaj, G.S.......................................................................56 Karube, Haruki; Katatani, Naoharu and Kitagawa, Kazuma ........................................................................56
Comparative studies on the genital and sub-genital abdominal segments of five species of dragonflies (Anisoptera: Odonata)........................................................................................... Allochthonous organic matter as a food resource for aquatic invertebrates in forested streams ..............................................................................................................................................
Thomas, Manu, Gunasekaran and Mohan, Daniel .........................................................................................57
Graa, Manuel A. S. ............................................................................................................................................57
POSTER PRESENTATIONS..................................................................................................... 59 1. Preliminary observations of reproductive behavior in Arigomphus villosipes (Anisoptera: Gomphidae)............................................................................................................... 2. Site fidelity, mating success and reproductive strategies in males of Libellula fulva (Odonata: Libellulidae) ................................................................................................................... 3. 4.
McMillan, Victoria E...........................................................................................................................................59
Nagy, H. Beta; Lszl, Zoltn; Szllassy, Nomi; Szkely, Annamria & Dvai, Gyrgy .......................60 Sathe, T.V.; Bhoje, P.M., Kavane, P., Shinde, Kiran; Yadav, R. P. & Pandharbale, A. R. ........................61 Schenk, Kamilla ...................................................................................................................................................61
Mating behaviour in damselfly Ischnura senegalensis (Rambur)........................................... Egg distribution, mate-guarding intensity and offspring conditions in dragonflies .....
5. Fat reserves and wing spot size, but not wing spot and body colour intensity, are related to male success during territorial contests in Hetaerina americana................................. 6. The effect of thermal pollution on dragonfly populations...............................................
Contreras-Garduo, Jorge; Buzatto, Bruno Najera, Karla & Crdoba-Aguilar, Alex ...............................62
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7. Evaluation of the odonata community in Galician rivers (NW Spain) that are affected by hydroelectric power stations...................................................................................... 8. Comparative molecular genetic analysis of the population structures and dynamics of two aeshnid species (Odonata: Aeshnidae) in Namibia........................................................
Mihokovic, Nino & Slavikovski, Ana ...............................................................................................................62
Rodrguez-Guntn, I.; Prez-Bilbao, A.; Gonzlez, A.; Alonso, A. & Garrido, J. ......................................63
9. Hybridisation and inheritance of female-limited colour polymorphism in two ischnurid damselfly species (Odonata)......................................................................................... 10. Genetic consequences of a man-made bottleneck in Orthetrum coerulescens: A microsatellite system to study fine scale population dynamics................................................. 11. The effect of larval density on female polychromatism and body size in Ischnura graellsii
Timm, Janne & Hadrys, Heike ..........................................................................................................................63
Van Gossum, Hans, Snchez-Guilln, Rosa Ana & Cordero Rivera, Adolfo ............................................64
Wargel, Antonia, Giere, Sandra & Hadrys, Heike...........................................................................................65
Gonzlez de Castro, Ins; Van Gossum, Hans & Cordero Rivera, Adolfo ................................................65
12. Preliminary results of a multi-year study of phenology and development of Anax junius in Maryland, U.S.A................................................................................................................
13. Relative frequency of Ischnura elegans and I. graellsii (Odonata: Coenagrionidae) in the Galician coast ............................................................................................................................
May, Michael L. ...................................................................................................................................................66
Snchez-Guilln, Rosa Ana & Cordero Rivera, Adolfo.................................................................................66
14. Present dragonflies in As Gndaras de Budio. Site of Community Importance (Nature 2000 Network) .................................................................................................................. 15.
Alonso, A.; Garrido, J.; Prez-Bilbao, A.; Gonzlez, A. & Rodrguez-Guntn, I. ......................................67 Ferreira, Snia & Grosso-Silva, Jos Manuel ..................................................................................................68
Present knowledge on the Odonata of Serra da Estrela Natural Park (Portugal)....
16. Odonata of continental Portugal: mapping the knowledge and identifying geographical gaps............................................................................................................................. 17. Dragonflies and damselflies of a montane tropical rainforest in Papua New Guinea 18.
Ferreira, Snia; Grosso-Silva, Jos Manuel; Soares-Vieira, Patrcia & Sousa, Pedro .................................68
Oppel, Steffen......................................................................................................................................................69 Sathe, T.V.; Shinde, Kiran; Bhoje, P.M., Thite, H.S. & Patil, R.G. ..............................................................70
Biodiversity of dragonflies (Odonata) from Western ghats of Maharashtra, India .
19. On the available data concerning the Odonata of Peneda-Gers National Park (NW Portugal).................................................................................................................................. 20. A preliminary analysis of odonate species richness in Galiza (NW Spain) ...............
Soares-Vieira, Patrcia, Grosso-Silva, Jos Manuel & Ferreira, Snia ..........................................................70 Azpilicueta, Mnica; Rey Ra, Carlos; Docampo Barrueco, Francisco; Rey Muiz, Xos Carlos & Cordero Rivera, Adolfo ......................................................................................................................................71
21. Contribution to knowledge of the biology of Onychogomphus costae Slys, 1885 (Odonata: Gomphidae) in southern Spain .................................................................................. 22. Pond water regime and competition as key factors in the presence and life history of two Lestes damselflies (Odonata: Lestidae) ................................................................ 23. Expansion of Crocothemis erythraea in Ukraine.................................................................
Cano-Villegas, Francisco J. & Ferreras-Romero, Manuel..............................................................................71
Torralba Burrial, Antonio & Ocharan, Francisco J. .......................................................................................72 Khrokalo, Lyudmyla & Matushkina, Natalia ...................................................................................................72
24. Sensitivity and economy of monitoring the environment of a large lake using adult dragonflies .............................................................................................................................. 25.
Kurauchi, Yohei & Ubukata, Hidenori ............................................................................................................73 Termaat, Tim, Ketelaar, Robert & de Vries, Henk.........................................................................................74
Flight peak trends for dragonflies from the Netherlands ............................................
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26. taxa
Morphological characteristics of odonate eggs and early instar larvae specific to
Watanabe, Yoko ............................................................................................................................... 74
27. Interannual change measurement of the Odonata community existing in a Mediterranean river which put up with the by-product of a heavy mining accident happened in April 1998 (river Guadiamar, Andalusia, southern Spain).................................. 28. Relationships between the concentrations of heavy metals in dragonfly larvae and Louro river sediments ............................................................................................................. 29. 30.
Ferreras-Romero, Manuel; Cano-Villegas, Francisco J. & Rubio-Soler, M. Isabel.....................................75
Rodrguez-Liares, G.; Garrido, J.; Bendicho, C.; Lavilla, I. .........................................................................76 Rodrguez-Liares, G.; Garrido, J.; Bendicho, C. & Lavilla, I.......................................................................77 Fernndez-Martnez, Miguel A..........................................................................................................................77
Evaluation of three species of dragonfly larvae as biomonitors of heavy metals ....
Dragonfly folklore in Galicia, Northwest Iberian Peninsula.......................................
31. A new genus of Coenagrionidae (Odonata, Zygoptera) from the Pantepui region of Venezuela, with descriptions of four new species. ................................................................ 32.
de Marmels, Jrg..................................................................................................................................................78 Mihokovic, Nino .................................................................................................................................................79
WAPODOKEY.................................................................................................................
33. A comparative study of haemocytes in male and female of the dragonfly Crocothemis servilia Drury ..................................................................................................................
Ashok, Jagtap; Shinde, Kiran & Sathe, T.V. ....................................................................................................79
Informal presentations ................................................................................................................... 80 Reflections on the 2003 International Odonatology Symposium, Beechworth, Australia...
Author index.................................................................................................................................... 81 LIST OF PARTICIPANTS .......................................................................................................... 84
Hawking, John H.................................................................................................................................................80
Abstracts Book 4th WDA International Symposium of Odonatology, Pontevedra 26-30 July 2005 Tuesday 26 July
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Oral contributions Forests as habitats for dragonflies (Odonata) Corbet, Philip S.

I.C.A.P., University of Edinburgh, Scotland, U.K.; present address: Crean Mill, St. Buryan, Cornwall TR196HA, U.K. <mail@pcorbet.vispa.com>
The ways in which forests can be inferred, or shown, to meet the habitat requirements of dragonflies are reviewed globally. The relationship between dragonflies and forests is examined along a latitude spectrum in the Northern Hemisphere, from the Arctic Circle to the equator, a transect along which species diversity progressively increases, and the microclimate within forest becomes steadily more permissive for occupancy by the several stages in the dragonfly life history. In mid-temperate latitudes dragonflies use forests mainly for aestivation as prereproductive adults, a strategy functionally similar to the siccatation exhibited by tropical dragonflies in seasonal-rainfall regions. Tropical rainforest is the planets most diverse terrestrial ecosystem, with regard to species and habitats. It provides habitats for many species of dragonflies, for some or all of their life- history stages. Many such species, and their behaviour and ecology, remain undescribed. For biologists, including odonatologists, the foremost challenge of our time is that this irreplaceable storehouse of biological information faces imminent threat of destruction before its contents can be placed on record.
Odonata in Bornean tropical rain forest formations: diversity, endemicity and implications for conservation management Orr, Albert G.
School of Australian Environmental Studies, Griffith University, Nathan, Q 4111, Australia. <agorr@bigpond.com>
The island of Borneo was originally almost completely covered by closed canopy tropical rainforest. Owing to an aseasonal, hot, perhumid climate and high rainfall, forests were well supplied with streams and standing water. Consequently the rich, largely endemic odonate fauna must have evolved in association with these forests, and non-forest species, common today in disturbed land, must formerly have been rare opportunists in forest gaps or localised lacustrine species. It is estimated that at least 70% of the fauna is presently confined to forest habitats and probably depends on forest for its survival. This study relates odonate distribution to a mosaic of complex tropical rain forest formations in Brunei. The tiny sultanate of Brunei still enjoys about 80% forest cover, representative of all the seven major formations found on the island and a great many of the 30+ subformations, and results from a nation-wide survey of odonates from most habitats are considered to be broadly applicable to the entire island of Borneo and many other parts of equatorial south-east Asia. Greatest odonate diversity, both alpha and beta, and greatest endemicity, is found in the primary lowland mixed dipterocarp forests, especially those growing in highly dissected landscapes such as occur at the KBFSC, at the edges of the central uplands. High diversity and endemicity is also found in swamp forest, especially freshwater swamp, with certain endangered peat swamp formations also important. The
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highly vulnerable Kerangas forest harbours fewer species, none uniquely, and the mangrove fauna is still more depauperate, with a single wide-ranging specialist. Secondary dipterocarp forest is certainly less rich in odonates than primary forest, but lack of sites for parallel comparisons makes it difficult at present to state how serious this effect is. These results emphasise the importance of conserving a wide range of primary forest formations to achieve satisfactory odonate conservation, a strategy congruent with the conservation of charismatic land-based vertebrates and forest peoples.
Threat levels to odonate assemblages from invasive alien tree canopies Samways, Michael J.
Department of Entomology and Centre for Agricultural Biodiversity, University of Stellenbosch, P/Bag X1, Matieland 7602, South Africa. <samways@sun ac za>
Dragonflies are well-known to be sensitive to light conditions, with the various species having a range of light conditions that they prefer. When these conditions are changed, such as by human removal of the tree canopy, the odonate assemblage changes accordingly, with forest species being replaced by species preferring sunlit habitats. Most of the South African species, including the national and local endemics, are mostly species that inhabit sunlit habitats, especially those fringed with indigenous grasses and bushes. During the 20th century, many of the South African riparian corridors became invaded and radically transformed by alien trees, especially Acacia spp. As these trees are a threat to hydrological processes, a massive national Working for Water Programme was started to clear riparian zones of these alien trees. These trees were also posing a major threat to local biodiversity, especially endemic odonates. Some species were even on the verge of extinction as a result of shading of their habitats by the alien trees and various synergistic impacts such as over-abstraction of water and damage to the banks by domestic livestock. The recovery of some of these odonate species as a direct result of alien tree removal has been absolutely remarkable, and is a strong message in support of genuinely effective and positive conservation action, involving removal of alien trees.
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Specialists vs. generalists among dragonflies - the importance of forest environments to form diverse species pools Sahln, Gran
Ecology and Environmental Sciences, Halmstad University, P. O. Box 823, SE-30118 Halmstad, Sweden. <goran.sahlen@set.hh.se>
In Scandinavia more species occur in forested environments than in agricultural areas. Some authors attribute the high number of forest species to extensive river and wetland networks. But since there are fewer species also in some agricultural areas with numerous wetlands, the explanation must be another. It is known that forestry affects species composition. Remove the trees, the environment changes and some species disappear. The time elapsed after forestry activity is important to species survival. While undisturbed forest habitats house the greatest number of species, partivoltine species decrease during the first 5-10 years after disturbance. Univoltine species are not affected - in fact the univoltine species present here are also part of the species pool of agricultural areas; they are true generalists. Analysing species composition in detail between lakes in different seral stages during forest regrowth gave ~90% separation between stages using discriminance analysis. Moreover, an even better separation was achieved analysing (semi)aquatic plant communities at the shoreline. Plants and odonates are connected, the species responding to the habitat's form and structure rather than to, e.g., acidity or nutrient levels. Forestry thus affects the very structures needed for survival. What kind of structures are we dealing with? A comparison between agricultural and forested areas, classifying species according to habitat preferences, showed that in treeless habitats there were fewer specialists as well as generalists in constructed wetlands, compared to older ponds and lakes. Also the latter habitats, however, had fewer species than the adjacent forested lakes. Looking at constructed wetlands <10 years of age showed that those close to forest habitats (even small clumps of trees) had, on average, more than twice as many breeding species than those in more open areas. Trees are obviously important to the species at least during some stage of their life. All species would probably live well in waters in open areas, but some will not survive unless a forest habitat is available at a moderate distance from their breeding waters. We are thus talking of larval as well as adult habitat, not forgetting egg-laying substrates. Forests thus seem to have what agricultural areas dont. To maintain a high diversity within a landscape, several seral stages and many different wetlands, surrounded by a diverse matrix of plants (including. trees) must be present. All this seems to lead to one general rule: Forests harbour specialists, while open landscapes are the playgrounds of generalists.
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Movement behaviors of odonates in heterogeneous forest landscapes Taylor, Philip D.

Acadia University, Patterson Hall, Rm. 323, Wolfville, Nova Scotia, B4P 2R6 Canada. <philip.taylor@acadiau.ca>
We have explored the population distribution and structure of the peatland dragonfly, Leucorrhinia hudsonica, in a forested landscape of western Newfoundland. Using surveys and mark-release-recapture experiments, we have shown that the amount of forest in the landscapes influences both the extent of movement among peatlands, and possibly the hydrology and pH within peatlands. In combination, these abiotic and biotic influences are associated with higher Leucorrhinia hudsonica abundances in landscapes containing less forest. Thus, the effects of forest harvest may influence abiotic factors influencing abundance at the scale of the individual peatland, and biotic factors at the scale of multiple peatlands. Precisely how such influences might translate into higher abundances are unknown, but are likely a result of altered processes at multiple and interacting scales.
The importance of forests to neotropical dragonflies Paulson, Dennis

Slater Museum of Natural History, University of Puget Sound, Tacoma, WA 98416, USA. <nettasmith@comcast.net>
Many species of Odonata depend on forests as habitat, not surprising as about two-thirds of the Earths land surface in subarctic latitudes was forested at the dawn of civilization. Of the 217 genera of Odonata in the New World, 175 (81%) include species that appear to be associated with forests. Forests are especially important in the tropics, as 88% of 164 Neotropical genera include forest-based species, while this is true for only 55% of 71 Nearctic genera. Forests may be less used at higher latitudes because of thermal considerations. Forests may be important to adult dragonflies by providing shade and humidity to avoid overheating and desiccation and by providing shelter from wind and predators. Few data are available to distinguish these alternatives, not necessarily mutually exclusive. Forests also provide breeding habitats, as there are many types of forested wetlands, especially in the tropics. In addition, forest conditions extend the life of temporary wetlands. Phytotelmata occur only in forests, and they furnish specialized breeding habitat for species in several families. Finally, forests are of great significance as seasonal or daily retreats for open-country species.
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Habitat use by pseudostigmatid damselflies: their future in fragmented forests Fincke, Ola M.
Department of Zoology, University of Oklahoma, Norman, OK 73019 USA. <fincke@ou.edu>
In the microhabitats of water-filled treeholes and bromeliads in neotropical forests, larval odonates, particularly those in the family Pseudostigmatidae, comprise the guild of top predators. Of these, the giant damselfly, Megaloprepus caerulatus, a monospecific genus which ranges from Mexico to Amazonia, seems to have a disproportionate effect on the population dynamics of its prey, which include mosquitoes, other odonates, and the tadpoles of several anuran genera. Within its geographic range, Megaloprepus is a reliable bioindicator of primary moist or wet forest. Both the larvae and adults are susceptible to drying conditions; the species is notably absent from tropical dry forests. Male Megaloprepus defend large tree holes where mating occurs. Hence, the presence of this species depends not only on a non-random assortment of tree species, but on trees above some minimum size. In a lowland moist forest of Panama, although adult Megaloprepus colonized tree holes in both old mature (> 400 yr-old) forest and contiguous young secondary forest (> 60 < 100 yrs), they were more abundant in primary forest, whereas two co-occurring Mecistogaster (linearis and ornata) were more common in secondary forest. Similarly, in a forest on the Atlantic slope of Costa Rica, Megaloprepus failed to colonize artificial holes adjacent to recently logged forests. The co-occurring M. linearis, colonized pots in both forest types. Mecistogster modestus, whose larvae develop in bromeliads found in both primary and secondary forests, was more abundant in secondary forests where the higher light environment favored a greater abundance of bromeliads. Collectively, these data suggest that Megaloprepus, and by analogy Microstigma rotundatum, which replaces Megaloprepus in some parts of Amazonia, may be more susceptible to forest fragmentation and than Mecistogaster and Pseudostigma, which can also occur in tropical dry forests. By releasing Megaloprepus from a boat and following their flight, it appeared that adults rarely cross large expanses of water (or open land). In a Mexican population, males lack the sexually dimorphic wing pattern characteristic of the more southern populations. Genetic analysis indicated population divergence sufficient for sub-species status. Although this divergence likely reflects a past geological barrier, current populations are threatened by rapid deforestation, resulting in severe habitat fragmentation. The characteristics which make Megaloprepus caerulatus a reliable indicator of primary wet forests appear to also make this species more susceptible to local extinction, relative to co-occurring pseudostigmatids.
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What matters to tropical forest dragonflies? African impressions Clausnitzer, Viola1 & Dijkstra, Klaas-Douwe "KD" B.2
1 2
Graefestr. 17, 06110 Halle, Germany <violacl@gmx.de> Gortestraat 11, NL-2311 MS Leiden, The Netherlands <dijkstra@nnm.nl>
Ecological traits of rainforest species and possible reasons for their disappearance when forest is opened up are discussed, based on examples from tropical Africa. We believe a combination of the low insolation in forest habitats and interspecific competition is a key factor segregating forest and openland species. While openland species cannot cope with low insolation inside the forest, forest species have evolved a tardy lifestyle to cope with the rainforest environment but are out-competed by more aggressive openland species once the forest is opened up. Most of our observations support this hypothesis, although the reality may be more complex, including additional factors such as water chemistry, habitat structure, substrate and availability of food to larvae. The majority of Africas openland dragonflies are widespread, whereas most forest species have a localised distribution. We speculate how climatic oscillations and associated largescale habitat shifts that Africa has experienced, and different dispersal capacities of the species, may have governed speciation across the forest-savannah ecotone. Phylogenetic reconstruction of groups that straddle this habitat divide, linked to ecological observations, may elucidate evolutionary reactions to landscape change. The reaction of odonate assemblages to forest loss is easily investigated in Africas imperilled forests. Because many of these forests are believed to be relatively young and highly forest-adapted species may have very low dispersal capacities, comparative ecological research of forest-dependent odonate assemblages in- and outside ancient forest refugia is recommended.
Historical and recent population genetics: Any news to tell us about the impact of fragmentation on afrotropical forest odonates? Hadrys, Heike
ITZ, Ecology and Evolution, TiHo University, Hannover, Germany, and Yale University, Dept. Ecology and Evolutionary Biology, OML, New Haven, CT 06520 USA <Heike.Hadrys@ecolevol.de>
Inferring historical microevolutionary and ecological processes based on present day patterns is a major challenge for evolutionary ecology and conservation biology. A recent scenario proposes that the tendency of species to retain their ecological niches over evolutionary time scales and their failure to adapt to new environmental conditions (phylogenetic niche conservatism) is a key factor in subdividing populations and initiating radiation processes. The forests of eastern Africa provide ample opportunities to study phylogenetic niche conservatism. They belong to the oldest, most threatened, heavily fragmented ecosystems and habour a high density of endemic species. A prime example to study the interplay of fragmentation and phylogenetic niche conservatism is the treehole breeding damselfly Coryphagrion grandis, an endemic of the Eastern Arc and Coastal Forests of Kenya and Tanzania. As member of the tree hole community its distribution depends on closed
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canope of moist forests. Treeholes are important forest freshwater habitats and their community is highly prone to adverse effects of fragmentation. The breeding habitats of C. grandis in Kenyas and Tanzanias remaining coastal forest patches, differ greatly in size, isolation and physical characteristics. Population genetic analyses focus on the question how historical and recent fragmentation events, habitat use and ecological conditions have affected population dynamics and speciation in C. grandis. A significant substructuring between distinct populations (forest fragments) in correlation to geographic distance was detected. The genetic diversity within populations was very low with no correlation to forest size. Recent phylogenetic and morphological studies have shown that C. grandis is the only member of the neotropical family of giant damselflies (Pseudostigmatidae) and a Gondwana relict. Consequently species radiation could have been expected, as this is the case for many insect species in this area. Instead evolutionary constrains in niche usage and the phylogeographic history have limited species radiation and left C. grandis as the only species of giant damselflies on the african continent. As an indicator species for primary forests and a relict of the Eastern Arc and Coastal Forests the population history of C. grandis may reflect the health and history of the remaining forest patches in a more general way.
The structure of the Coenagrion mercuriale populations in the New Forest, southern England Thompson, David J. & Watts, Phillip C.
School of Biological Sciences, The Biosciences Building, Crown Street, University of Liverpool, Liverpool L69 7ZB, UK. <D.J.Thompson@liverpool.ac.uk>
The damselfly Coenagrion mercuriale (Charpentier) is a poor disperser and susceptible to habitat fragmentation/loss and is protected by European legislation. An analysis of Capture-Mark-Recapture (CMR) data indicate that the population network on Beaulieu Heath in the New Forest comprises some 40,000 individuals. A nationwide genetic study indicates that the New Forest is a principal reservoir of genetic diversity for UK C. mercuriale. The New Forest is, however, presently best characterised as five genetic units. Several small, isolated populations of C. mercuriale in the New Forest show substantial genetic differentiation from the principal populations on Beaulieu Heath, Setley Plain and Mill Lawn Brook. Isolation is bought about by preventing dispersal across intervening areas of unsuitable habitat such as forest, farmland or road. Although habitat loss is a principal concern for the persistence of this species, the pattern of limited movement to proximate sites highlights the need for a network of suitable of habitat patched for this species' persistence and to slow the rate of genetic erosion at peripheral sites.
Abstracts Book 4th WDA International Symposium of Odonatology, Pontevedra 26-30 July 2005 Wednesday 27 July
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Mate location and competition for mates in relation to sunflecks of forest floors Watanabe, Mamoru
Graduate School of Life and Environmental Sciences, University of Tsukuba, Ibaraki 305-8572, Japan. <watanabe@kankyo.envr.tsukuba.ac.jp>
Although most species have a maiden flight away from water and sexually immature adults stay in the forests foraging for food, mature males tend to remain in the forests in some species, such as the white-legged damselfly, the emerald damselfly, opportunists of the hyaline-winged males of Mnais damselfly and so on. To locate females in the forests, males mainly perch in sunflecks (a sunlit site in the forest floor) and adopt sit-and-wait tactic. Some of them try to occupy the perching site. Territorial behaviour of males of the whitelegged damselfly, Platycnemis echigoana, was described at sunflecks in climax deciduous forests. They showed the patrolling flight along the periphery of the sunfleck, and hovering above the perching site, suggesting that such flight was a display of the occupation of sunfleck. Flight behaviour of the emerald damselfly, Lestes sponsa, in the forest floor also showed male-male interference in accordance with the mating behaviour, and a lek-like system was discussed. Solitary males interfered in copulation in the forest floor, while some solitary males were also observed on the shoreline of the pond throughout the day, but they did not harass the pairs ovipositing in tandem. Although hyalinewinged males of Mnais pruinosa costalis adopt sneak tactics, a male that failed in occupying a perching site to intercept females entering the territory is called an 'opportunists', which moves around forest floor with sunflecks to search females. From the viewpoint of sperm displacement, the longest copula duration observed in the opportunists was advantageous. These observations point to functional relationships with habitat selection and thermoregulation. Perching behaviour under direct sunlight at sunflecks was shown to result in considerable variation in thermoregulatory properties. The relationships of thermoregulation to mate location strategy are different among species. A percher under direct sunlight will be benefit to approach a passer-by quickly, and to fight with other males. Forest structures with sunflecks were discussed in viewpoint of the habitat selection acting on the female choice. These relationships also have relevance to other behaviours, particularly oviposition behaviour by water. Adults that showed mating behaviour in the forests oviposit in tandem by water. The importance of sunflecks in the forest floor was discussed in relation to the life history strategy of the damselfly species inhabiting forests.
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Lack of innate recognition of species or morph identity in Enallagma damselflies Fincke, Ola M.
Department of Zoology, University of Oklahoma, Norman, OK 73019 USA.<fincke@ou.edu>
In several families of Zygoptera and Anisoptera, female-specific color polymorphisms seem to be a female response to sexual harassment by mate-searching males. A suite of male mimicry hypotheses assume that, relative to the heteromorph, the male-like andromorph is invariably more difficult for males to recognize as a potential mate. In contrast, the learned mate recognition (LMR) hypothesis predicts that males learn to recognize both morph types and hence lack an innate preference for either morph. In an experiment with the damselfly, Enallagma civile, males raised to maturity with only other males were less likely to react sexually to females of either morph, compared to males reared with either morph. Moreover, when the nave males did react sexually, they displayed no preference for either morph type, in contrast with control males, which were significantly more likely to react sexually to the morph female with which they had been reared. Similarly, in a field population of Enallagma ebrium that had never experienced individuals of their sister species, E. hageni, E. ebrium males readily attempted tandems with E. hageni females of both morph types, although none achieved tandem due to the incompatibility of male claspers and the females' mesostigmal plates. These results, which suggest that closely related Enallagma males learn not only morph but species identity, hold important implications for the rapid speciation known to have occurred within this North American clade.
Large-scale variation in female morph frequencies of the polychromatic damselfly Nehalennia irene Van Gossum, Hans1,2; Beirinckx, Kirsten1; Forbes, Mark1 & Sherratt, Thomas1
Department of Biology, Carleton University, 1125 Colonel By Drive, Ottawa, Ontario, K1S 5B6, Canada Evolutionary Biology Group, University of Antwerp, Groenenborgerlaan 171, B-2020 Antwerp, Belgium. <Hans.VanGossum@ua.ac.be>
1 2
Females of the Sedge Sprite damselfly Nehalennia irene occur in two distinct morphs: the andromorph and the gynomorph. Spectacularly, we show impressive geographic variation in female morph frequencies with consistently high gynomorphic frequencies in one region, while in a different region consistently high andromorphic frequencies were observed. Also, excitingly we inform for the first time on within-population temporal (seasonal and among years) variation in female morph frequencies. More specifically, we report on differences in female morph frequencies in populations at variable distances (2 km 5000 km within Canada) and in nearby populations at different times (2 weeks 12 years within Ontario). In all of the populations sampled in British Columbia (n = 9) over 90% of females were andromorphs, whereas populations sampled in central (n = 20) and eastern Canada (n = 10) comprised on average about 10% andromorphs. Analysis of female morph frequencies at 5 different populations in Ontario throughout a summer gave clear evidence of within-season changes in morph frequencies, as well as differences in andromorph frequencies among nearby locations. N. irene sampled in the same eight locations 12 years apart showed dramatic changes in their population structure. In
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particular, the andromorph frequencies at these ponds in the two decades were not significantly correlated, while male densities at these times were negatively correlated. The one factor that co-varied with andromorph frequency in the majority of our spatiotemporal analyses was male density, and we discuss the significance of this relationship in the light of contemporary explanations of female-limited polymorphism.
Documenting odonate behaviour by drawing from films Hilfert-Rppell, Dagmar

An der Wasserdurche 32, 38162 Cremlingen, Germany. <rueppell-film@t-online.de>
Drawings of odonates from memory are seldom true. Getting images from the field and use them as model is the only suitable method to draw odonate behaviour. Even to photograph odonates is very difficult and only one moment is available from still photos. Better results promises filming. Video-filming is a first attempt. With this you get 25 images per second. By means of a short speed shutter a selected choice of behavioural images could be the result. But the images are of poor quality and lack most of the wing movements, because of the low temporal resolution of video-filming. Odonates recognize each other by a high temporal resolution so they might see each other in the same way as we do in slow motion films. These films offer a very good opportunity to draw even special and very fast behavioural patterns. In my talk I show different examples with different filming methods and the results as drawings. Most of the examples originate from our recent book on European Calopterygids, such as fighting males, emerging females, landings, take-offs, alternative reproductive behaviour or predation. But even with slow motion films details of the body or of the wings are often hidden, so morphological studies should be done in addition.
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Ovipositor and egg-laying behaviour of Odonata: phylogenetic implications Matushkina, Natalia

Dept. Zoology, Fac. Biology, Kiev National University, pr. Glushkova 2, b.12, K680 Kiev, Ukraine. <matushkina@list.ru>
Odonata is an insect group with various types of oviposition. Deposition of the eggs endophytically, i.e. inside of plants, was considered as the most ancient mode of egg laying. However, morphological peculiarities of endophytic ovipositor and peculiar properties of the egg sets in endophytic egg clutches have not been considered in some phylogenetic reconstructions, probably because of their expected uniformity. However, our recent comparative morphological studies have revealed significant varieties of the morphology of the ovipositor, muscular pattern and egg-laying behaviour within some zygopteran and anisopteran families. This study aimed to test the phylogenetic value of some ovipositionrelated characters in Odonata. 29 species from 27 genera representing 13 families of recent suborders of Odonata, were included in the data matrix. These were coded for 75 characters, 42 of which concern skeleton, 27 muscular and 6 behavioral peculiarities. Since the nearest winged insects group, Ephemeroptera, lacks any ovipositor structures, the apterygote Pedetontus sp. (Microcorrhyphia, Machilidae) having well-developed ovipositor, was selected as an outgroup. All the oviposition approaches known for dragon- and damselflies were considered, but peculiarities of the endophytic oviposition were emphasized. The separate coding (i.e. the presence and the shape of the character scored as separate characters) was used. Missing data and inapplicables were coded as question marks (reductive coding). Cladistic analysis was performed with PAUP* 4.0b10. Heuristic searches were carried out under simple parsimony, performing TBR branch swapping, random addition sequence with 10000 replicates and unlimited maximum number of MAXTREES. Successive approximations weighting was used with the maximum value of the retention index as its the weighting function. The resultant consensus trees demonstrate: Zygoptera, Anisoptera and Anisozygoptera are represented by separate clades, each supported by at least one synapomorphy; Anisoptera (Epiophlebia) have 4 synapomorphies. Monophyly of the family Aeshnidae is supported by 2 synapomorphies. Lestidae are represented by a separate clade with the most strongly supported monophyly among Zygoptera. The highest phylogenetic value is inherent to the following studied characters: (1) external morphology: form and hairiness of the stylus, structure and position of the supporting edge; (2) internal morphology: form of the posterior apophysis, some features of the muscles 5, 6, 7 and 8; (3) behavior: pattern of the eggsdeposition in endophytical clutches. Also, distribution of some morphological and behavioral characters within Odonata is traced and possible scenarios of their evolution are discussed.
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Flashes in flight communication between odonate males Rppell, Georg

An der Wasserfurche 32 38162 Destedt, Germany. <rueppell-film@t-online.de>
Do odonates with coloured wings fly differently? In slow motion films three libellulids from Namibia show different flight styles. Circling flight and wingstandstills enhance the signalling effect of the coloured wings. In Calopterygids and Megaloprepus coerulatus (Pseudostigmatidae) the wingbeat pattern has changed from the normal zygoptercounterstroking to parallel-stroking.This doubles the colour areas seen at any one moment. Calopteryx splendens and C. virgo- males do interact, when displaying threatening flight, by showing their blue wings, while females only accept the male of its own species, showing the special courting flight. This is species-specific, characterized by the different phaserelationships of the both wingpairs. The Japanese Mnais-species with orange wings, beat them in the same way as the European species and even the male-forms with hyaline wings do so. The flight of odonates with coloured wings is so special that I propose to call it:signalling fligt. The talk includes slow-motion films from Namibia, Japan, Europe and Panama. Pre- and postmating mechanisms of reproductive isolation between Ischnura graellsii and I. elegans (Odonata: Coenagrionidae) Snchez-Guilln, Rosa Ana & Cordero Rivera, Adolfo
Grupo de Ecoloxa Evolutiva e da Conservacin, Departamento de Ecoloxa e Bioloxa Animal, Universidade de Vigo, EUET Forestal, 36005 Pontevedra, Spain. <rguillen@uvigo.es>
The hybridization process provides an excellent opportunity for the study of microevolutionary phenomena. In this paper we test the importance of isolation barriers between two sibling species of the genus Ischnura (I. elegans and I. graellsii) using laboratory experiments. Both species are morphologically and genetically very similar and are sympatric along part of Galician coast. Even if heterospecific matings seem uncommon in odonates, previous results indicate that when both species coincide in the same locality, hybrid matings are commonly observed. Among the prezygotic barriers we studied the incidence of temporal segregation, the ethological and mechanical isolation mechanisms (incompatibility for tandem formation and for starting copulation), the prevention of fertilization (gametic or genetic incompatibility, conspecific sperm procedence, fertility and fecundity). Postzygotic barriers like hybrid inviability (longevity, mortality, etc), hybrid sterility (fertility and fecundity) and the hybrid breakdown were also examined. Three hybrid generations were reared in the laboratory under controlled conditions. Results indicate that prezygotic barriers are more effective than postzygotic mechanisms to prevent hybridization: mechanical problems make 90% of mating attempts between male I. graellsii and female of I. elegans unsuccessful, while gametic incompatibility is negligible. However, when the mating direction is between male I. elegans and female I. graellsii the mechanical barriers are not very effective, only preventing 10% of mating attemps. Nevertheless, 60% of these females do not lay eggs and if they lay, only 56% of them are fertile. We conclude that I. graellsii is disappearing due to genetic absortion by I. elegans, because it is the female of graellsii the only involved in heterospecific matings.
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Does ovaries composition vary between species with different mate-guarding intensities? Schenk, Kamilla
Institut fr Geokologie, Technische Universitt Braunschweig, Langer Kamp 19c; D-38102 Braunschweig, Germany. <k.schenk@tu-bs.de>
Past experiments (Schenk, K., Suhling, F. & Martens, A. 2004. Egg distribution, mateguarding intensity, and offspring characteristics in dragonflies (Odonata). Animal Behaviour, 68, 599-606) suggest that there are two different types of egg clutches in libellulids: Egg size decreases significantly during oviposition in species which perform non-contact guarding during oviposition. In contrast, in species ovipositing in tandem, egg size is randomly distributed. The different egg clutch patterns may also be explained by morphological differences between the females genitalia. However, although there are some studies on morphology of ovaries, none of these compared morphology with the function of the ovaries nor could they explain the different egg size distributions within the clutches. In this study I proposed that the ovarios differ between the two guarding types. Therefore, microtom sections of the ovaries of thirteen libellulid species were done and several egg and follicle cell parameters were measured. Dragonfly prey capture: Vision, decision, and flight Olberg, Robert 1 and Worthington, Andrea2
1 2
Dept of Biology, Union College, Schenectady, NY, 12308, USA Dept of Biology, Siena College, Loudonville, NY , 12211, USA. <WORTHINGTON@siena.edu>
Dragonflies employ their excellent visual systems to intercept other insects in flight. Using high-speed video, we recorded the behavior of the libellulid dragonfly, Erythemis simplicicollis, a perching predator. We captured adult, foraging females and placed them in a large flight cage. By the following day, the dragonflies foraged actively from perches within the cage. We videotaped individual dragonflies taking off in pursuit of a 2 mm white bead moved on a fine wire, simulating passing prey items. We analyzed 130 clips at 500 frames per second, digitizing the coordinates of the head position and angle, thorax, abdomen and the bead location before and during the interception flights. We used these data to address three questions: (1) How does the perched dragonfly estimate distance to a passing object? (2) How does the dragonfly compute the direction for takeoff? (3) How is the target fixated visually during flight? Before takeoff, head movements kept the image of the moving bead on or near visual midline and also on or near the high acuity dorsal fovea about 55 above the horizon. Rapid, saccade-like head movements moved the image across the fovea, a behavior that could provide motion parallax information for distance estimation. The angle at which the dragonfly launched could be accurately predicted from a calculation of the angular velocity of the bead 40 ms before takeoff. During the flight the dragonflys head rotated relative to the thorax to fixate the bead on visual midline and on the dorsal fovea. These head movements were much faster that the corrective steering of the thorax by the wings. We are currently looking for neural pathways that provide proprioceptive information about head angle to the wings to realign the body with the head during flight.
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Copulatory behaviour in hybrid matings between Calopteryx haemorroidalis and C. splendens Cordero Rivera, Adolfo
Grupo de Ecoloxa Evolutiva e da Conservacin, Departamento de Ecoloxa e Bioloxa Animal, EUET Forestal, Universidade de Vigo, Campus Universitario, 36005 Pontevedra, Spain. <adolfo.cordero@uvigo.es>
The reproductive behaviour of odonates is under intense selection due to sperm competition and female choice. Typically, copulation starts with a phase where males use their genitalia to remove sperm from previous mates (stage I), and only at the end of copulation insemination takes place (stage II). Previous experiments with zygopterans in laboratory conditions showed that copulation duration with mated females is almost twice as long as with virgins in species whose females have a spermatheca, but this behaviour was probably not the result of sperm competition, because males could not remove sperm from the spermatheca in the studied species. In Calopteryx, males can or cannot remove sperm from the spermatheca, depending on the relative size of male and female genitalia. In some populations males are able to physically remove sperm, but in others they cannot introduce their penis horns inside the narrow spermathecal ducts. I tested the influence of this phenomenon on copulatory behaviour of C. haemorrhoidalis and C. splendens using an outdoor insectary and the hand-pairing technique, in Pontecorvo (Central Italy). In the studied population of C. haemorrhoidalis male genitalia is narrower than the spermathecal duct, and can physically remove sperm. This is unknown for C. splendens. Copulation lasted 2.040.30 min (N=22) in C. haemorrhoidalis, 2.010.15 (36) in C. splendens, and 1.930.18 (12) in hybrid matings between male splendens and female haemorrhoidalis. Nevertheless, when males of haemorrhoidalis were had-paired to females of splendens, copulation lasted only 1.200.14 (9). The reduction in copulation duration is due to a shorter stage I, and stage II remains unchanged. Assuming that males control copulation duration, this suggests that males of haemorrhoidalis are able to easily remove sperm from females of splendens, but not in the opposite direction. In the field, spontaneous copulations have been observed between male splendens and female haemorrhoidalis; these females sometimes lay eggs after the hybrid mating and a putative hybrid has been found, suggesting that hybrid fertilization takes place.
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Sperm ejection as a cryptic female choice mechanism in odonates Crdoba-Aguilar, Alex

Instituto de Ecologa, UNAM, Apdo. Postal 70-275, Coyoacn, Mxico, D. F. Mxico. <acordoba@ecologia.unam.mx>
The few odonate studies of sperm use suggest that females spend apparently more sperm than what seems necessary during oviposition (sometimes, females may have their sperm stores reduced to a 50% after a single oviposition episode). Furthermore, some studies have documented that females eject sperm during and after copulation. This posits the question of whether sperm reduction may be interpreted as a cryptic female choice mechanism. Using two zygopterans (Ischnura denticollis Burmeister and Enallagma praevarum Hagen) and one anisopteran (Pantala flavescens Fabricius) for different purposes, I investigated whether: a) females mate multiple and with different males; b) sperm reduction and ejection take place; c) there is inter-individual female variation in sperm reduction; d) female sperm reduction variation and number of eggs laid is consistent across pairs of females mated to the same male; and, e) the ejected sperm comes from the vaginal duct which may obstruct the egg passage. Results indicate that females: a) mate multiply; b) show a heavy reduction in stored sperm but this takes place prior to oviposition; c) ejected sperm; d) vary inter-individually in sperm reduction which is inversely related to the number of eggs laid; e) when mated to the same male showed similar reductions in sperm stores and egg load; and, f) only rarely, the vaginal duct had sperm. This suggests that a heavy sperm reduction is common in this insect order and is not explained by an excess of sperm in the vaginal duct that may obstruct the egg passage. I suggest that females sperm shortage is better explained as a cryptic female choice mechanism aimed to favour the sperm of some males. This study provides exciting research avenues for future studies of female choice in an animal taxa whose sexual biology has been otherwise regarded as male controlled.
Survival and mating success of American Rubyspots in relation to body size (Odonata: Calopterygidae) Serrano-Meneses, M. A.; Szkely, T. & Crdoba-Aguilar, Alex
Department of Biology and Biochemistry, 4 South, Claverton Down, University of Bath, BA2 7AY, Bath, United Kingdom. <bspmasm@bath.ac.uk> Instituto de Ecologa, Universidad Nacional Autnoma de Mxico, 04510, Mexico D.F.
1
Sexual differences in body size are widespread amongst animals. The explanations for the evolution and maintenance of sexual size dimorphism (SSD) are numerous (i.e. natural selection, sexual selection and fertility selection), and they have been tested in several taxa by observational and comparative studies. Odonates exhibit an exceptional variation in SSD, ranging from reversed dimorphic species (females larger than males) to species where males are larger than females. Despite a growing number of studies, there is still disagreement as to what causes males and females to differ in size across this insect order. First, in a single-species study we investigate the effects of body size on survival and mating success in a Mexican population of the American Rubyspot (Hetaerina americana), a sexually dimorphic species in which males are larger than females. In this species, males exhibit
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large red spots at the base of each wing that are the result of male-male interactions. Males defend territories along streams and rivers, and mating success is determined by the ownership of a territory. Preliminary results show that large males hold territories for longer periods than small males, sustain longer territorial fights, obtain more copulations than non-territorial males, mate with larger than average females and show higher proportions of wing pigmentation. These results suggest that in this territorial damselfly species, large male body size is selectively advantageous since it is related to territory holding potential and mating success. However, further analyses are required to test whether body size influences survival. Second, I will present results of a comparative study in which I investigate the roles of sexual- and fertility selection on SSD in Odonates by using phylogenetic comparative methods.
Differences in survival of mated and unmated males of Libellula fulva: a four year study Szllassy, Nomi1; Szab, Zoltan D.2 & Nagy, H. Beta3
Dept. of Science Education, Babes-Bolyai University, Cluj, Romania Dept. of Systematics and Ecology, Babes-Bolyai University, Cluj, Romania 3 Dept. of Hydrobiology, Debrecen University, Debrecen, Hungary
1 2
We studied the recapture and survival rate of a nearly closed population of Libellula fulva along a small creek in East-Hungary. Our goal was to make a comparison between the survival rate of mated and unmated males. During a four year study we marked a number of totaly 672 males on their right wing. Through the study period (May and June) we observed the daily mating activity of marked individuals between 9.00h and 16.00h walking along a 500 m section of the stream. The mark-resight data of mated and unmated males were analysed with the MARK software. Survival and resighting probability were estimated separately, using capture-recapture models. In 2000 the survival rate did not differ between the two male groups: mated (observed at least once in wheel-position) and unmated (not involved in pairing during the observation period) and it was constant over time. In 2001 the survival rate of mated individuals was higher than the unmated ones, but the difference is not significant. Mated males had significantly greater forewings and hindwings than unmated ones. The probability of recapture was higher in the case of mated males and varied with time in both years. The number of hours spent with observation did not explain the variation in the recapture rate. In 2002 and 2003 it was a slightly difference between the survival rate of two male groups. In the last two years the recapture rate varied again with time. In 2002 the selected model showed group interaction, in 2003 the recapture rate of two male groups showed no differences.
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Female reproductive behaviours of different Chlorocyphid species in the OrientalAustralian region Gnther, Andr
TU Bergakademie Freiberg, IOEZ, AG Biologie/kologie, Leipziger Strasse 29, D-09599 Freiberg, Germany. <andre.guenther@ioez.tu-freiberg.de>
The Chlorocyphidae are well known for their mostly complex male aggressive and courtship behaviour. In the result of a comparative study of several Chlorocyphid species in the islands of Sulawesi, Bali and New Guinea and in the Malay Peninsula a wide range of female reproductive behaviour could be observed as well. The females of most of the species choose among territory-holding males for mate. The courtship behaviour of the males seems to be less important than the quality of the oviposition site and the female experience at the same site during the previous days. Most of the species show well developed refusal behaviour for avoiding male interference with oviposition. The female tactics vary from classical behaviour like spreading of the wings and escape by rapid flight to interspecific behaviour e.g. in syntopic populations of Heliocypha perforata and Libellago semiopaca. This type of behaviour has been observed the first time for the two species.
The global dragonfly assessment Spector, Sacha1 & Naskrecki, Piotr2

Center for Biodiversity and Conservation, American Museum of Natural History, New York, NY. <spector@amnh.org> 2 Conservation International, Museum of Comparative Zoology, Cambridge, MA
1
While freshwater ecosystems cover only 0.8% of the Earths surface, they are the habitat of virtually 100% of the worlds 5,600 species of dragonflies and damselflies. Rapid degradation or loss of wetlands and freshwaters are resulting in the the imperilment of large numbers of dragonfly and damselfly species. Regional analyses have suggested that greater than 20% of dragonfly/damselfly species may be endangered or threatened, and there is a growing number of presumed extinct taxa. In order to develop a strategic plan to effectively conserve dragonfly diversity and, more broadly, freshwater biodiversity, the American Museum of Natural History and Conservation International have initiated a global assessment of the conservation status of the worlds dragonfly and damselfly species. This effort represents the first effort to comprehensively document the distribution, population status, and conservation needs of any invertebrate taxon at the species level. Conservation status of the worlds dragonfly/damselfly species are being assessed within the IUCN SSC Red List Program framework, on the basis of: a) documented range sizes/trends; b) known population sizes/trends. Data on the distribution of species is being derived from museum collection specimen records, regional or private databases of collection records, and locality information contained within the historical literature. Where possible, population trends are being assessed from modern monitoring data and/or the historical patterns of collection records. These distribution and poulation trend data will be coupled with additional expert information on threats, population trends, etc. for each
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species to produce final assessments. The GDA will identify the most threatened species and critical habitats for this group of invertebrates, as well as lead to the inclusion of dragonflies and damselflies in other global initiatives such as CIs Hotspot analyses and the Alliance for Zero Extictions map of priority conservation sites.
Why are endemic Odonates endangered in oceanic islands Ogasawara? Karube, Haruki
Kanagawa Prefectural Museum of Natural History, 499, Iryuda, Odawara, 250-0031, Japan. <qtmfc457@ybb.ne.jp>
The Ogasawara Islands (situated about 1000 km south of Tokyo, Japan) are oceanic islands consisting of two main inhabited islands (Chichi-jima and Haha-jima) and a group of about 10 other small satellite islands which are now uninhabited. There are 5 known endemic dragonfly species: Boninagrinon ezoin, Indolestes boninensis, Rhinocypha ogasawarensis, Hemicordulia ogasawarensis, Boninthemis insularis. These islands have 300 years of history of human impact, but most of the endemic dragonflies were common prior to 1980. During the 1980s these dragonflies started to become regionally extinct in the northern part of Chichi-jima. The extinction rapidly expanded to the south after 10 years, and in the 2000s they have become completely extinct. Also in Haha-jima, the same extinction process began from the mid 1990s, and after 10 years it changed to a situation similar to Chichi-jima. Now there is only one species, R. ogasawarensis remaining. At first the reason for this surprising rapid decrease was thought to be due to large typhoon damage or insecticides sprayed for harmful insects. However in my research, the former idea was rejected because of the existence of a large population of dragonflies in the small islands near Chichi-jima and Haha-jima, and the latter is not true. This regional extinction in both the main islands happened not only to Odonata, but also to many other kinds of insects, such as butterflies, beetles and wasps. Finally, it has become clear that the insect extinction has been caused by the introduced lizard Green Anolis from North America. This lizard was accidentally introduced during allied occupation period after WWII. It is a strong insect predator, and in Chichi-jima, it was introduced at the end of the 1960s from Guam. The distributional expansion coincides exactly with the decrease of dragonflies. The lizard population is now estimated to be about 500,000-4,000,000. In Haha-jima, the introduction happened in the 1980s and expansion occurred more quickly. The insect extinction is confirmed to be limited to these two main islands (the Green Anolis islands). In the other satellite islands the dragonfly population has remained constant. Now, I. boninensis is only known to exist in one small island, Otouto-jima (5 km2). H. ogasawarensis is only known to exist in two islands: Ani-jima (8 km2) and Otouto-jima. Both populations are very small and estimates are under 250 individuals. The status of further three species is not as drastic, but they are also endangered. Now, their situation has become weak, so, if for example strong drought happened, they would easily become extinct. I believe this is the first report in the world suggesting that an invasive introduced predator is the cause of extinction among dragonfly species.
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Dragonflies of riverine habitats : assessment as indicators of biodiversity and environmental integrity. Riservato, Elisa & Bogliani, Giuseppe
Universit degli Studi di Pavia, Dipartimento di Biologia Animale, Laboratorio di Eco Etologia, Piazza Botta 9, 27100 Pavia, Italia. <elisa.riservato@unipv.it>
We investigated the potential use of Odonata coenosis in the assessment of biodiversity and environmental integrity in riverine habitats. Surveys were carried out in different habitats within the Ticino Natural Park (Northern Italy), where species-habitat preferences where investigated. Forty sampling stations were studied, ten in each of the following 4 different habitats: 1 - springs with effluent canals; 2 - the main river banks; 3 - agricultural canals; 4 - oxbow lakes and gravel pit lakes. Microhabitat characterization - In each sampling station we carried out standardized samplings and recorded larval and adult habitat characteristics. Macrohabitat characterization - For each sampling station we produced a GIS based habitat map of surrounding landscape, trough which we looked for ecological preferences at the landscape level. Larval ecology. In each station, larvae where collected using a standardized procedure; percentage composition of micro habitats was described, and chemical and physical parameters of water were measured. Adults ecology. A survey grid was created in order to analyze the ecological preferences of adult dragonflies. The base transect was 8 m x 20 m, divided into twenty 2 m X 2 m squares. Transects length was established with the aim of contacting as much as possible microhabitat diversity, and hopefully the majority of species. Transects of different length were used: 20 m in springs with effluent canals; 60 m in oxbow lakes, gravel pit lakes and agricultural canals; 100 m in the main river banks. All transects were located along the shore line and in each square the position and type (species, sex and behavior) of every individual was mapped. In each square, habitatl composition was described according to definite classes. Every station had been surveyed 5 times, during two months. Habitat preferences at both the microhabitat and the landscape scale of different species are discussed.
Spatial heterogeneity of the dragonfly assemblages in the landscape scale: assessments using newly created small ponds as traps in the catchment area of lake Kasumigaura Kadoya, Taku; Suda, Shin-ichi, Washitani, Izumi & Tsubaki, Yoshitaka
Department of Ecosystem Studies, Institute of Agriculture and Life Science, The University of Tokyo, Bunkyo-ku, Tokyo 1138657, Japan. <kadoya@e-mail.jp>
Definition of an appropriate sampling unit is a critical matter to assess the distributional heterogeneity of species at large scale successfully. Since adult lentic dragonflies have concentrating distributions at ponds in a landscape in their reproductive periods, we may be able to use a whole pond as a sampling trap to assess the distributional patterns of dragonfly species in a landscape.
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In the catchment area of Lake Kasumigaura (36N, 140E) in Ibaraki Prefecture, eastern Japan, more than 100 small man-made ponds so called dragonfly ponds were created by a NGO leading a lake ecosystem restoration project during 1999 2002 to provide suitable habitats for dragonflies and refugia for endangered aquatic plants. In such newly created ponds, colonization by some lentic dragonfly species will occur. If immigration is the dominant process determining the dragonfly community structures (i.e., species composition and their abundances), those in the ponds will reflect the abundances of species around the ponds, and assemblages of sites in geographical proximity will tend to support more similar assemblages than sites that are further apart. In order to test the hypothesis, we conducted adult dragonfly censuses on 49 dragonfly ponds during the period of adult dragonfly activity in 2003 and analyzed the spatial patterns of community structures among ponds. We recorded a total of 1087 dragonflies of 18 species (6 zygopterans and 12 anisopterans). The variances of community structures of adult dragonfly species were summarized using principal component analysis (PCA), and the spatial component of the site scores was modeled using a third-ordered polynomial of the ponds coordinates (x1: longitude, x2: latitude). To separate the effect of environmental differences among ponds, the environmental factors which have significant effects on adult dragonfly species habitat preferences such as pond size, vegetation cover and open sky ratio were included in the model as covariables. We obtained a significant regression model that explained spatial patters of community structures of adult dragonfly species among ponds. This result indicates that it is possible to use dragonfly ponds to describe spatial heterogeneity of the dragonfly assemblages in the landscape scale.
Fecundity and fertility of parthenogenetic Ischnura hastata Lorenzo Carballa, Olalla & Cordero Rivera, Adolfo
Grupo de Ecoloxa Evolutiva e da Conservacin, Departamento de Ecoloxa e Bioloxa Animal, EUET Forestal, Universidade de Vigo, Campus Universitario, 36005 Pontevedra, Spain. <olalla.lorenzo@uvigo.es>
Recent theories of sexual selection stress the importance of conflicts over reproduction in shaping the reproductive traits of males and females. When the reproductive interests of both sexes coincide, which only occurs under strict monogamy, sexual conflict is absent. But in the majority of cases, there is a conflict of interests between sexes over the number of matings and reproductive decisions. It has been suggested that males are selected to "harm" females if this increases male reproductive success, even at the expenses of female fitness. One prediction of such hypothesis is that sperm is selected to maximize the probability of fertilization, and this sometimes can cause a decrease of fertility due to multispermy, genetic incompatility, toxic seminal products that harm females, and so on. Populations of Ischnura hastata at the Azores are parthenogenetic, while this same species has sexual populations in North America. This system is therefore adequate to test theories of sexual conflict, because we expect harmful effects of sexual conflict to disappear in the parthenogenetic populations. We have measured fecundity and fertility in 209 parthenogenetic females over eight laboratory generations. Furthermore, we obtained egg clutches from 14 females collected in sexual populations in Mexico, and estimated fertility in their first clutch laid in captivity. Parthenogenetic females (N=26) of the first generation (that emerged in the laboratory from field collected larvae) laid an average of 235 eggs over
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their lifetime with a fertility rate of 0.9250.017. Females from Mexico laid 62 eggs, of which 0.8660.052 were fertile. This difference is significant (GLM with binomial errors, deviance ratio=9.77, p=0.003). Fecundity of parthenogenetic females was lower in generations that emerged during winter. Fertility was different between generations but was not lower during winter. These results suggest that fertility is higher in parthenogenetic individuals, which is in agreement with sexual conflict predictions.
Habitat selection and Egg production in Sympetrum infuscatum females living in a forest-paddy field complex Susa, Koichi & Watanabe, Mamoru
Graduate School of Life and Environmental Sciences, University of Tsukuba, Ibaraki 305-8572, Japan. <susa974@hotmail.com>
Although larval habitats of Sympetrum infuscatum are rice paddy fields, all adults leave the rice paddy fields for forest gaps after emergence, and stay there during their sexually immature stages. In late summer when they matured, some of them visit the rice paddy fields in tandem flight for oviposition. However, the result of mark and recapture samplings showed that a half of males and more than four fifth of females in the population remained perching in the forest gaps, where no mating behaviour was observed. To evaluate the habitat selection of S. infuscatum females in the forest gaps, we examined the fecundity by dissection. In the early morning (0600-0900) when most individuals rested in the forest gaps, there were two kinds of females in the viewpoint of the number of mature eggs loaded: females that loaded more than 400 mature eggs seemed to be willing to oviposit on that day, and that loaded less than 100 mature eggs. In the morning (0900-1200) when oviposition activity was high in the rice paddy fields, females that remained in the forest gaps loaded about 100 mature eggs. The mean number of mature eggs loaded in the females in the rice paddy fields was significantly higher than that in the forest gaps. However, the variance was large because we captured both females that had started to oviposit and that had just terminated to oviposit. In the afternoon (1200-1500), the mean number of mature eggs in females remained in the forest gaps increased. In the evening (1500-1800), there were wide variety in the number of mature eggs loaded in the females, probably because of females that had released most of their eggs had returned to the forest gaps and because of females that remained in the forest gaps had developed their eggs up to nearly 500. In the artificially ovipositing experiment, females in the rice paddy fields released eggs significantly faster (53 eggs/min) than females in the forest gaps (13 eggs/min). When the females were kept in the cage feeding 3 times per day, they developed 120 mature eggs per day. Consequently, females may have to remain in the forest gaps for more than 4 days to develop their eggs up to 500 mature eggs, the number which might allow females to go to the rice paddy fields for oviposition.
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Ecology of Odonata inhabiting permanent Namibian desert springs Martens, Andreas1 & Suhling, Frank2
1Abteilung Biologie, Pdagogische Hochschule, Bismarckstrasse 10, D-76133 Karlsruhe, Germany. <martens@ph-karlsruhe.de> 2Institut fr Geokologie, Technische Universitt Braunschweig, Langer Kamp 19c, D-38106 Braunschweig, Germany
Natural perennial surface water in the interior parts of Namibia only occurs at widely separated springs around mountains. These waters host a very diverse and unique Odonata assemblage, which is threatened due to the habitat restriction of several species, as well as by recent habitat loss and degradation. Species occuring permanently at these waters, including Pseudagrion kersteni, Crocothemis sanguinolenta or Orthetrum julia, differ significantly in seasonality, dispersal and lifetime characteristics from species colonising temporary waters. These characteristics are important to understand why opportunistic colonisers with rapid development -which are dominant elsewhere- do not outcompete them at those special habitats.
Abstracts Book 4th WDA International Symposium of Odonatology, Pontevedra 26-30 July 2005 Friday 29 July
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Artificial parthenogenesis in Aeshna nigroflava Martin Watanabe, Yoko

4-14, Nishida-cho, Nishinomiya-shi, Hyogo, 662-0034, Japan. <mer-yoko.watanabe@nifty.com>
We reported the case of artificial parthenogenesis in Stylurus oculatus in 1997 and 1999, and proceeded to examine other odonate species we could obtain. While parthenogenesis could not be observed in most of the species examined, it was detected in Aeshna nigroflava Martin. As an egg of this species, which belongs to the Aeshnidae, is large in size and its embryo colors conspicuous at the time of its embryonic development, some abnormal cases of development were observable. Some picture images of such cases will be presented. Aeshna nigroflava oviposits mainly in September. Eggs soon start developing, then go into diapause in the pre-inversion state in order to pass the winter. Normaly-developed eggs hatched, but the percentage was small. Most of the eggs collected in the Kansai region started development, though the percentage of cases of malformation was quite high, while an extremely small percentage of eggs collected in the Hokkaido region began developing in one sample, only one egg developed. We did not raise the larvae after hatching. In the case of Stylurus oculatus, also, there was a wide individual variation, many cases of malformation were observed, and there were many which did not grow even though they hatched.
Distribution and overlap of ranges of Lestes parvidens and Lestes viridis in southeastern Europe (Odonata: Lestidae) Weihrauch, Florian1; Olias, Marko2; Bedjani, Matja3; Marinov, Milen4 & alamun, Ali5
1 Jgerstrae
21A, D-85283 Wolnzach, Germany. <Florian.Weihrauch@t-online.de> 29, D-09599 Freiberg, Germany; 3 Kolodvorska 21/b, SI-2310 Slovenska Bistrica, Slovenia; 4 P.O. Box 134, BG-1000 Sofia, Bulgaria; 5 Center za kartografijo favne in flore, Podrunica Ljubljana, Zemljemerska 10, SI-1000 Ljubljana, Slovenia
2 Humboldtstrae
Bartenev (1910) and Morton (1922) were the first to recognize Lestes viridis specimens from southern Russia and western Turkey, respectively, which differed clearly from central European specimens in the shape of male appendices. Later this taxon was described as Lestes viridis parvidens by Artobolevskii (1929), based on specimens from Crimea, Ukraine. However, as L. parvidens then was regarded as an southeastern vicariant of the nominotypical European race, European odonatologists did not take notice of it for almost 60 years until G. Lehmann collected specimens clearly pertaining to L. parvidens near Ravenna, Italy (Schneider 1986). Only in the 1990s the occurrence of L. parvidens in Italy and Greece received more attention (Lohmann 1993), and detailed studies by C. Utzeri and co-workers (e.g., Utzeri et al. 1994) presented evidence for the specific rank of the taxon and for a wide distribution of both L. parvidens and L. viridis within peninsular Italy. However, their actual ranges in the Balkans hitherto remained unclear as all published records from that region prior to the mid-1990s did not differentiate between the two spp. and thus are unusable.
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To get more detailed information on the actual distribution of L. parvidens and L. viridis in southeastern Europe, we tried to compile a sufficient amount of data to address this topic. These data include all published records with clear differentiation between the two spp., a number of specimens from zoological collections checked by ourselves, and some unpublished records we took in the past years. The mapping of these data presents the following distributional patterns: The range of L. parvidens extends from the East across the Balkans, almost reaching Austria in Northeastern Slovenia, covering the entire Istrian peninsula and peninsular Italy south of River Po, and reaching Corsica in the west. On the other hand, the range of L. viridis extends to the Black Sea in southeastern Bulgaria and the Aegean Sea on the Chalkidiki peninsula, and covers entire Italy to southern Sicily. No records of L. parvidens are available from Sardinia and Sicily. However, we traced records of possible hybrid populations from Sicily as well as from peninsular Italy, Slovenia, Croatia, and Montenegro.
Odonata larvae and drought in Australia: Ecological development for life in an unpredictable climate Hawking, John H.
Murray Darling Freshwater Research Centre, Cooperative Research Centre for Freshwater Ecology, P.O. Box 991, Wodonga, 3690 Australia. <john.hawking@csiro.au>
Australia is one of the driest continents resulting in a fauna which has evolved morphologically and adapted ecologically to withstand periods drought (without free water). To survive periods of drought dragonflies generally have a diapause stage in their development stage, either in the egg or larvae. In the northern hemisphere diapause in the egg stage is common; however, it is currently not recorded in Australian Odonata. Although there is some evidence though that it may occur in some species of Hemiphlebiidae and Lestidae. In Australian dragonflies, the drought resistant stage occurs in the larvae, the longest stage of the life cycle, from 1 -10 years. This stage is most likely to be subjected to drought conditions. The adaptation to drought appears to be mainly in the more ancient Australian dragonflies of the families, Megapodogrionidae, Synthemistidae, Aeshnidae (Telephlebiidae), Corduliidae (Austrocorduliidae, Pseudocordulidae). The larvae of many Megapodogrionidae species inhabit swamp areas which all dry in summer, surviving buried in the muddy sediments. The larvae of Podopteryx selysi (Megapodogrionidae) lives in water filled tree holes in rainforests jungle. Some Telephlebiidae and Pseudocorduliidae live a semi aquatic life feeding on terrestrial invertebrates. The Petaluridae dig deep burrows, through which the larvae move between the water and terrestrial environment. Many of the Synthemistidae and Gomphomacrominae larvae inhabit streams which dry in the summer and survive by burrowing into the sediment under large rocks or into the mud. These species are absent from the very arid regions of Australia and tend to be confined to the higher rainfall areas of the eastern and western ranges. These species experience extremes in the rainfall patterns causing long periods of drought and floods. Adaptation by these ancient species, through time has allowed them to cope successfully to the unpredictable climate by having a long larval life cycle, adapting morphologically and developing drought resistant ecological strategies.
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The influence of environment and phylogeny on the determination of morphological, behavioural and life history traits in dragonfly larvae Suhling, Frank1 & Sahln, Gran2
Department of Environmental System Analysis, Institut fr Geokologie, Technische Universitt Braunschweig, Langer Kamp 19c, D-38102 Braunschweig, Germany. <f.suhling@tu-bs.de> 2 Ecology and Environmental Science, Halmstad University, P. O. Box 823, SE-30118 Halmstad, Sweden
1
Freshwater communities can be classified along different gradients, e.g. with respect to the extents of disturbance or drying. Identifying and examining traits that influence the distribution of species along such gradients is crucial to the understanding of community structure. Theory predicts that traits should differ between species that live in temporary and permanent waters because of differing major environmental variables, which are drying and predator presence, respectively. Since resources and energy available are limited, traits of organisms associated with a certain part of the continuum cannot be maximised simultaneously, and therefore trade-offs with respect to traits associated with each kind of habitat can be expected. Species, however, will also be influenced by their evolutionary history, i.e. the traits of their common ancestors. We studied morphological and behavioural traits associated to foraging and life history traits of libellulids co-occurring in the Namib desert. We aim to analyse which traits depend on phylogenetic inertia or adaptive radiation, i.e. on the type of habitat. We tested for effects of the habitat type (temporary and perennial water) and phylogeny by analysing one species of each of the three dragonfly genera, Crocothemis, Orthetrum and Trithemis, from one of the habitat types. We found that behavioural traits did not vary between the habitat types but between the genera. By contrast, life history traits varied with habitat, but not with genus. Of the morphological traits some were influenced by habitat and others by phylogeny. Interestingsly, some traits of the same organs, e.g. of the eyes may vary with the habitat, while others do so with phylogeny. We aim to interpret the adaptive values of variations in morphology, behaviour and life history.
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Quaternary environmental change along the Western Escarpment of Africa and the distribution of Namibian Odonata Marais, Eugene
National Museum of Namibia, P.O. Box 1203, Windhoek, Namibia. <insects@natmus.cul.na>
The Western Escarpment of Africa, stretching from the South Africa's Cape region to central Angola, has experienced considerable environmental changes during the late Quaternary. Though the environmental history of this south-western part of Africa was considered to have approximated desert conditions for a long time, recent advances in palaeoenvironmental research question that assumption. Results from different locations along the escarpment suggest that conditions during the Last Glacial Maximum (LGM) and the Holocene were highly variable on a millennial scale, cautioning against palaeoenvironmental models that view the LGM as a uniform cold and arid phase. LGM vegetation seems to have differed markedly from that of the Holocene, indicating more humid conditions in the highlands along and above the escarpment. More mesic conditions along the arid to extremely arid coastal plains are therefore likely, even though drier conditions dominated in the interior center of Southern Africa. Results from geomorphological studies of ephemeral river-courses on the coastal plain confirm more regular fluvial discharge regime than at present. Such palaeoenvironmental reconstruction may explain disjunct distributions in current populations of Namibian Odonata, with particular reference to rare species and associated genetic evidence that these species experienced significant bottlenecks in the past. Since environmental conditions are not only determined by rainfall, but is the result of complex interactions between atmospheric circulation (dominant wind, moisture sources), vegetation cover, solar radiation and temperatures, topography and other elements, the implication is that dragonfly migratory pathways and ecological prerequisites may have changed significantly within the past 8000 years. The Odonata would have responded in different ways to such changes, but those species that have been isolated as a result of climatic change contribute most information towards understanding the consequences of climate change. An environmental change from milder to more severe conditions in recent geological time may also explain the extraordinary level of endemicity in various other faunistic groups along the Namibian escarpment, while the patterns observed in the Odonata are invaluable for explaining or confirming the likely evolutionary pathways for endemicity.
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The occurrence of Cordulegaster sp. in Czech Republic result of influence of habitat ecological factors in different biogeographical regions? Holua, Otakar
Mendel University of Agriculture and Forestry, Department of forest botany, dendrology and geobiocenology, Zemdlsk 3, CZ - 613 00 Brno, Czech Republic. <holusao@email.cz>
The CZ-territory (dimension 78 864 km2; altitudes from 117 to 1602 m a.s.l.) is divided into biogeographical subprovinces - Hercynian, Westcarpathian, Polonic and Northpannonic. Hercynian region builds up by metamorphic rocks and deep vulcanites. Relief is tectonic breakdown flush surface. Westcarpathian region (in the CZ-territory) is composed by sediments of flysch (sandstones, claystones, shales). The high superelevation is typical feature. Polonic region intervenes only marginally from north a few stabilized and floppy sediments and they were modelled by Pleistocene glacier. Pannonic region is a valley, which is built up by unconsolidated sediments loesses, calcareous sands and sediments in broad river floodplains. During 1992-2004 was carried out intensive investigation of watercourses in all forestry areas in CZ. Two species of genus Cordulegaster - C. bidentata and C. boltonii occur in the CZterritory. C. bidentata was found first of all in the Westcarpathian region in altitude 300-950 m a.s.l. (centre 400-500 m a.s.l.). Spring areas and very small brooks are habitats of occurrence. Egg-laying was always observed in spring areas. Habitats of larvae: depth of water column 1-10 cm, width of watercourse 12-110 cm, volume flow 0,01-2 l.s-1, bottom material is mixture of mud and soft grit (gravel) (sometimes in calcareous springs) geests of sandstones and claystones; high density of larvae - 10 larvae on 1 m2 (5 larvae of last instar). Abundance of population changes with the changes of parent rocks (from SW towards to NE) the highest in Bl Karpaty Mts. and Vsetnsk vrchy hills geests of close-grained sandstones (often clacareous), claystones; lower in Moravskoslezsk Beskydy Mts. poorer (siliceous) sandstones, and the lowest in Silesian Beskydy Mts. ragstones and pudding stones. In Hercynian part is occurring only in the eastern part - geests of greywacke. C. boltonii was found above all in the uplands in Hercynian region in altitude 200-900 m n.m. (centre 600-700 m a.s.l.). Meandering brooks and lesser rivers are habitats of occurrence. Egg-laying was observed in small brooks. Habitats of larvae: depth of water column 20-100 cm, width of watercourse 0,75-5 m, volume flow 30-50 l.s-1, bottom material is coarse-grained sand to brash with stones geests of gneisses, mica schists. The highest abundance was found in region of rsk vrchy hills and Jihlavsk vrchy hills. Different occurrence is given by geomorphology of regions and thereby of watercourses. But according to results the occurrence of Cordulegaster sp. is determinated by geological differences, first of all by geests of parent rock for occurrence of larvae.
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Notes on Crimean Odonata (Crimea, Ukraine) Khrokalo, Lyudmyla1 & Prokopov, G.2
1 Kyiv National Taras Shevchenko University, Volodymyrska str. 64, Kyiv, Ukraine 01033. <lkhrokalo@mail.ru> 2 Vernadskiy Tavricheskiy National University, Yaltynska str. 4, Simferopol, Crimea, Ukraine 95007. <prokopov@crimea.com>
Odonatological investigations in Crimea Peninsula have a long history. Early species descriptions are in Selys-Longchamps (1853) and Hagen (1856). The first report of 13 species from Simferopol environs was given by Brauner (1902), followed by other studies (Brauner, 1903; Pliginsky, 1913; Artobolevsky, 1915; Bartenev, 1912, b, 1915, 1919; Shorygin 1926). In a subsequent paper (Artobolevsky, 1929), 37 species were reported, and a new subspecies Lestes viridis parvidens (now: Chalcolestes parvidens Art.) was described from Crimea. Odonata were later used in entomological and hydrobiological assemblage studies (Tseeb, 1947; Maltsev, 1953; Temirova et al., 1984; Kiseleva, Vasyuta, 1984; Kiseleva, Jezernitsky, 1985; Kiseleva, 1992; Kiseleva, Vershytsky, 1998; Prokopov, 2001; Pyshkin et al., 2003; Prokopov, 2003). Thus, 48 species are known from literature, although four species (Coenagrion ornatum, C. lunulatum, C. armatum, Aeshna juncea) have been identificated by hydrobiologists on larvae only, suggesting the need for some confirmation. We have discovered 29 species from ad hoc collections of adults, exuviae and larvae from 1999-2004 in South and West coastal and montane parts of Crimea. Four species (Anax ephippiger, Orthetrum albistylum, Sympetrum flaveolum, S. danae) are recorded from the Crimea peninsula for the first time. An analysis of habitats of larvae has been made. Ten species were found in rivers, 7 in ponds, 7 in small lakes (including montane lakes), and one species in brooks and springs. The problems of identification of Calopteryx taurica larvae and distribution of Orthetrum anceps will also be discussed.
Biogeography and habitat affinity of the odonata fauna of SE Madagascar Schtte, Kai
University of Hamburg, Dept. Animal Ecology and Conservation, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany. <Brancsikia@web.de>
The Odonata of Madagascar have not been studied in almost any aspects of ecology, biogeography or conservation. Systematic work and inventories have been equally rare in the last decades. The current work is located in SE Madagascar where the last remnants of littoral forests in a heavily degraded ecosystem are found. Deforestation by the local human population is still in progress and deposits of heavy mineral sands are located in this area, even underneath the forest fragments, swamps and marshlands. QIT Madagascar Minerals will mine and rehabilitate immediately afterwards. For postmining rehabilitation efforts, conservation zones are established in some of the least degraded fragments. To measure results of rehabilitation, specific tools are necessary. In this region a drastic climatic change is caused by the Anosy mountain chain that is a weather barrier. The research was conducted from February to April 2004 and from October 2004 to June 2005 and preliminary results are presented. Basic inventories of Odonata were carried out in selected littoral, transitional and mountain forest fragments with water sources and the surrounding
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marshland and anthropogenic influenced areas. The sampled forest fragments were separated into different degradation levels. Species are roughly separated into forest and openland species and, based on this, into species that have the tendency to enter degraded forests. Species distribution in relation to habitat quality and the rainfall gradient was sampled. Both larvae and adult Odonata were collected and observed. At least one new species was collected. Larvae are reared to adults and genetic tissues of both were preserved to link them to the adults. Larvae will be described. With this and the knowledge of physical habitat quality, adult odonata as well as larvae can be suitable biological indicators that could be used as monitoring tools to measure rehabilitation success after the mining process.
Population differences in sexual selection intensity and immune response in two contrasting forest environments in the damselfly Hetaerina americana (Zygoptera: Calopterygidae) Contreras-Garduo, Jorge & Crdoba-Aguilar, Alex
Differences in life history traits may be a result of the specific adaptation processes faced by each population in a particular place. One trait which has received considerable attention in life history theory is immune ability. Recent results, for example, suggest that the machinery behind immunity may demand sustantial resources to the organism frequently at the expense of other vital functions. However, compromises between immunity and these other vital functions are expected to vary among populations in relation to the environment of each population. In this work we looked for differences in immune ability in two populations with contrasting environments (a semi-desertic, dry forest and a semi-tropical, humid forest) using the zygopteran Hetaerina americana. In this species, males bear a red wing spot which is related to male territorial and mating success: males with larger wing spots are more successful. Furthermore, our previous work has suggested that males show a compromise in expression between wing spot size and immunity (based on melanin encapsulation of a non-pathogenic agent) in which only males with larger wing spots are better at mounting immune responses. In this work we compared the following male variables between populations: (1) sex ratio, (2) copulation rate, (3) contest rate (4) figthing time, (5) immune ability; (6) time to complete the immune response; (7) repeatability of immune response during onthogeny, and (8) immune response after a costly territorial contest. We found differences between populations in all these variables which indicated that the semi-desertic forest population had a less intense sexual selection pressure (a more balanced sex ratio, higher copulation number per male, lower contest rate and fighting rate) than that of the semi-tropical forest. The former population was also able to mount a less intense immune response but exhibited smaller wing spots. Our interpretation is that due to the more relaxing selective regime, this population is not demanded to show the trend as intensively as that of the semi-tropical forest. However, there is a role for the environment as indicated by additional data as the semi-tropical forest seems to provide more food and a more benign temperature which can help to maintain a good immune response.
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Southern dragonflies expanding in Wallonia (South Belgium) : a consequence of global warming? Goffart, Ph.1, 2; Fichefet, V.2, 1; de Schaetzen R.3; Baugne J-Y.1; Lebrun, Ph.2 & Dufrne, M.1
1 Observatoire de la Faune, de la Flore et des Habitats (OFFH), Centre de Recherche Nature, Forts et Bois (CRNFB), Gembloux, Belgium. <P.Goffart@mrw.wallonie.be> 2 Unit dcologie et de Biogographie, Universit Catholique de Louvain, Louvain-la-Neuve, Belgium.; 3 Gomphus, Arquennes, Belgium.
The frequency of seven southern Odonata species has been watched in Wallonia over the last two decennia (from 1981 to 2000). They have clearly expanded in the meantime and this pattern is still highly significant when the data are corrected for the increase of prospection efforts and for the spatio-temporal heterogeneity of sampling. Moreover, reproduction evidences have been collected recently (from 1993 onwards) for all these species. Three distinct hypotheses are examined and discussed as possible explanation of the observed expansion pattern: (1) global warming, (2) change in aquatic habitats (eutrophication,...), and (3) intrinsic population dynamics. The rise of temperatures appears to be the main factor explaining the observed expansions.
Acknowledgments: all the collaborators of the Dragonfly Working Group Gomphus for data collecting, the Rgion wallonne (MRW/DGRNE) for funding the Monitoring Scheme.
The Dutch Dragonfly Monitoring Scheme: results and trends. Bouwman, Jaap1; Groenendijk, Dick1 & Plate, Calijn2
1 Dutch Butterfly Conservation, PO Box 506, NL-6700 AM, Wageningen, The Netherlands. <jaap.bouwman@Vlinderstichting.nl> 2 Statistics Netherlands, PO Box 4000, NL-2270 JM, Voorburg, The Netherlands
After a probationary year the Dutch Dragonfly Monitoring Scheme started from 1998 onwards and the number of monitoring plots increased quickly to nearly 400 plots counted yearly. About 40% of the plots are single species sites, which are counted at least three times a year during the flight-period of occurrence of the counted species. At all other sites, all dragonfly species are counted nine times a year. Counting is usually done by volunteers between May and September using a standardised method. Results are used to calculate yearly indices for each species, using the computer programme TRIM (Trends and Indices for Monitoring Schemes). This programme was developed by Statistics Netherlands for the analysis of time series of repeated counts with missing observations. One of the products of the Dutch Dragonfly Monitoring Scheme is the assessment of the total year-count for each particular year and species. The calculated indices can be used for overall evaluation of Dutch nature policy and conservation, and in the protection of endangered species like Leucorrhinia pectoralis and Aeshna viridis. The indices now show which species are declining, for instant the rare Leucorrhina pectoralis and Coenagrion hastulatum but can also show an increase for species like Calopteryx virgo and Libellula fulva. It is concluded that the Dutch Monitoring Scheme is a powerful tool in the conservation and protection of dragonflies and their habitats in the Netherlands.
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Dragonfly monitoring in northrhine Westfalia, Germany Conze, K.-J.

Listerstr. 13, D-45147 Essen, Germany. Klaus-Juergen.Conze@t-online.de
In northrhine-westfalia the current monitoring of dragonflies is done by the "Arbeitskreis Libellen NRW". This honorary working group for the protection and recording of dragonflies in NRW was founded in 1996. It has meanwhile established as a circle of about 200 persons and institutions all over NRW with at least 50 active collaborators. The work and the progress is organized by an inner circle of 15 colleagues. The database currently contains circa 20,000 reports, and this number is still growing fast since much of the old data, in addition to new observations every year, need to be entered. The AK organizes an annual meeting for all collaborators to report new information, and to give the opportunity to exchange experiences with each other. Circular letters and mailings also bring the latest information to every member. A weekend meeting is organized for this summer about observing and reporting dragonflies in a changing area (in 2005 it will be in Essen!). Very important, especially for instant information is the homepage: www.ak-libellen-nrw.de, where al lot of information is available. There it is possible to read the latest news of interesting dragonfly reports or download the recording manual and report sheet, the current red list, a bibliography or new dates. The AK also organize the training of colleagues for exuviae and adult dragonflies. It supports dissertations and other work on dragonflies. If anybody has problems to determine a special exuviae, he can send it to an experienced member, who does the determination (for further details please visit the homepage!). There is also an exhibition which can be rented out for presention at a requested public location in order to promote public relations for dragonflies. The AK is working in an efficient network together with the LBF (the major governmental institution for ecology and nature conservation in NRW), the Biologische Stationen (institutions for nature conservation on the level of districts), the GdO (the Association of German-Speaking "Odonatologists" - mean people who are interested in dragonflies) and other field-working groups (i.e. the AK Herpetofauna NRW). There are a lot of interesting results, some of them clearly related to climatic effects. So especially the outspreading of mediterranean species could be proved.
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When south goes north: mediterranean Odonata conquer Flanders (North Belgium) De Knijf, Geert & Anselin, Anny
Institute of Nature Conservation, Kliniekstraat 25, B-1070 Brussels, Belgium. <Geert.deknijf@inbo.be>; <anny.anselin@inbo.be>
Flanders lies in Nortwest-Europe, situated somewhat halfway between the Mediterranean region and Fennoscandia. This position is also reflected by its fauna and flora with several typical species with a Westsiberian or boreo-alpine distribution and also some with a more southern, mediterranean distribution. This is also the case for several Odonata species. The data are derived from the Dragonfly Working Groups database, containing > 55.000 records collected by more than 300 field collaborators. Species with their main distribution area in the Mediterranean part of Europe and which occur in Flanders are: Lestes barbarus, Coenagrion scitulum, Erythromma lindenii, Erythromma viridulum, Aeshna affinis, Anax parthenope, Orthetrum brunneum, Crocothemis eythraea, Sympetrum fonscolombii and Sympetrum meridionale. We analysed the data in different time periods: <1900, 1900-1949 and from then onwards per decade: 1950-1959, 1980-1989 and finally 1990-2004. To have an idea of the recent (>1980) evolution we made yearly analysis. We calculate for those 10 species the combined number of 5km grids for each period together with their relative weight. Before 1900 they were found in more than 20% of the investigated grids. From 1900 onwards it was less than 10% with an exception in the sixtees (15%). In the eighties it was again >20% and rocket since 1990 to nearly 60%! A similar view gives us the number of records. Since 1980, a first influx was noticed in 1983 and 1984. A second one started from 1989 and is still going on with a peak in 1994 and 1995. A peak was also noticed in 1999, 2000 and 2003. Erythromma lindenii and E. viridulum have always been present in Flanders, be it in low numbers. The first to establish themselves in Flanders were Lestes barbarus (1984) and Crocothemis erythraea (1989). They were followed by S. fonscolombii (1992), A. affinis (1994), C. scitulum (2003), S. meridionale (2003) and finally O. brunneum (2004). Some of them have become now very common and occur in 20% of the grids (Lestes barbarus and Crocothemis erythraea) or even 50% (Erythromma viridulum). In the 19th century, 8 of those 10 species were observed and some, e.g. Coenagrion scitulum and Sympetrum meridionale had dixit Selys (1866) at least temporary populations. Our data together with Selys saying suggest that many of those southern species were then present in Flanders, disappeard nearly completely in the period 1900-1980 and are now conquering Flanders.
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Towards an atlas of European odonates Kalkman, Vincent

Darwinweg 2, 2300 RA, Leiden. <Kalkman@naturalis.nnm.nl>
In the last decade a large number of regional and national distributional atlases have been published in Europe. As a result of this it has become possible to make dot-maps of the European distribution of odonates based on actually distributional data. Distributional atlases on European scale have been made for several other groups like mammals, birds and butterflies. All these projects have encountered numerous difficulties but have become a success in the end. The Dutch Invertebrate Survey has decided to sponsor the work on a European atlas by devoting a part of its office time for the organisation and work on the database. A project for a European atlas of dragonflies can only be a success when a large number of people and organisation will cooperate. The year 2005 will therefore be used to explore if a project for an atlas will find the broad support it needs. A proposal for the project will be outlined during the presentation and a number of examples of maps will be presented.
Abstracts Book 4th WDA International Symposium of Odonatology, Pontevedra 26-30 July 2005 Saturday 30 July
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Mapping potential habitats using environmental surrogate measures: Importance of forests for dragonflies in Japan Tsubaki, Yoshitaka
National Institute for Environmental Studies, Tsukuba, 305-8506 Japan. <tsubaki@nies.go.jp>
The success of biological conservation is measured in terms of the amount, quality and health of the biological diversity maintained (Samways 2005). Therefore, we need sound taxonomy and good distributional data, which are often not available for insects. Conservation area selection then depends on using coarse, but not inaccurate, data that are available on relatively well known species (or species assemblages) such as birds, butterflies and dragonflies. Therefore surrogate data are often used for making informed conservation decisions. Environmental surrogate measures include vegetation types, land systems or classes, climate and environmental domains. These environmental surrogate measures are often available from national geographic information facilities. Some countries, including Japan and Britain, have national recording schemes. Records reported by the network of volunteer recorders provided, to some extent, comprehensive coverage of the country. These are immensely valuable for determining how well or not species are doing over time, as well as the extent of the geographical ranges of species. The outcome has been the production of an atlas (Japan Integrated Biodiversity Information System), which provides an immediate visual overview of present geographical ranges. However, there are shortcomings with these record point maps. Firstly, the records are accumulated in an ad hoc manner, resulting in geographically biased records (for example, due to uneven spatial distribution of recorders). Secondly the data only record what is present and not what is absent. Thirdly, records are taxonomically biased because recorders may be keen to record rare species rather than common ones. Fourthly, abundance is neglected though it gives important survival implications for populations. I have developed a method for data quality validation, to overcome some of these shortcomings inherent to the national recording schemes. I have selected only high quality data from the original data set, for the use in the analysis of relational spatial database. As the results, I could detect a continuous array from an extreme generalist to a specialist in odonate fauna, which reflects species forest independency or dependency.
Genetic consequences of habitat specialisation and cryptic speciation in the genus Trithemis Giere, Sandra & Hadrys, Heike
ITZ Ecology & Evolution, TiHo Hannover, Bnteweg 17d, 30559 Hannover, Germany. <sandra.giere@ecolevol.de>
A comparative study of the genetic consequences of habitat specialisation in three African dragonfly species of the genus Trithemis discovered a new cryptic Trithemis species in the Okavango region in Namibia. Cryptic species are identical in morphology but genetically different. We studied and compared the population dynamics of T. kirbyi, T. arteriosa and T. stictica between different population sites in Namibia using two sequence markers (the mitochondrial ND1-locus and the 16S-rRNA region). The three species differs in their habitat requirements along a gradient from generalist (T. kirbyi) to specialist (T. stictica).
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Data analysis shows significant differences in the genetic diversity patterns between the populations and the species which correlates with the degree of the habitat specialisation. In contrast to the widely distributed T. kirbyi, T. arteriosa shows a lower sequence diversity but significant genetic substructures between the populations. The riverine T. stictica shows the lowest intra population and highest inter population sequence diversity. Here the sequence divergence (48 nucleotide substitutions/ 480 bp of ND1) between the two Namibian populations (Naukluft / Okavango) match those for separate species. However the first taxonomic characterization identified the Okavango specimens as T. stictica. Interestingly, the Namibia/Naukluft population is genetically identical to populations analysed from localities in Kenya, Tanzania and South Africa. Phylogenetic 16S rRNA analyses of the Naukluft/Okavango populations including eight more Trithemis species confirmed the presence of a cryptic Trithemis spec. nov. The speciation process will be discussed in the context of a possible habitat shift.
We acknowledge support from the BMBF (BIOTA Africa project S08)
The effects of Climatic Changes for the distribution of dragonflies in Europe and their possible effects on the biocoenosis of the waters Ott, Jrgen
L.U.P.O.GmbH, Friedhofstrasse 28, D-67705 Trippsadt, Germany and University of Landau, Fac.of Environmental Sciences, Im Fort 7, D- Landau, Germany. <L.U.P.O.GmbH@t-online.de>
In this paper first of all a general overview on the actual changes in the distribution of dragonfly and damselfly species in Europe will be given. Since the first records of a northward expansion of some species in the eigthies of the last century meanwhile a broad spectrum of southern species could be found expanding their range to the north and in higher altitudes: mediterranean species to northern Europe and recently also African species to southern Europe. This lead in the beginning to an increase of the darogonfly diversity and long-term monitoring programmes - if available - showed a clear change in the total dragonfly fauna of an investigated area. But recently also negative effects in some areas were registered, in particular a decrease of mooreland species after the extreme climatic situation of 2003. Several stenoecious species seem to be at risk and endangered under these conditions and the abiotic conditions may lead to a complete change of the coenosis. Examples for these changes are given and scenaria are presented for a future development of the European dragonfly fauna.
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Optimization of environmental monitoring schedule using adult dragonflies Ubukata, Hidenori and Sakoda, Tetsuo
Hokkaido University of Education, Kushiro, Hokkaido, 085-8580 Japan. <ubukata@kus.hokkyodai.ac.jp>
Using adult dragonflies to monitor freshwater ecosystems has advantages that enough amount of data are easily gathered and that information on rare species can be obtained, while the method has disadvantages, being susceptible to the weather as well as the time of day and of year, and corresponding less directly to the quality of the water than that using larvae or exuviae. The latter problem being dealt with in another presentation by Kurauchi and Ubukata, the former problem is challenged in this presentation: in designing a monitoring scheme using adult dragonflies, the optimal interval of periodical censuses that minimize the cost while the purpose of the monitoring is simultaneously satisfied should be sought. We made intensive route censuses of adult dragonflies on sixteen belts (each 50m long and 2m width) at Onnenai, Kushiro Marsh, Hokkaido, Japan, from June through October 2003. An index, 'species confirmation ratio' (SCR), was defined as the ratio of the number of species confirmed in a census to that confirmed by three censuses executed in that month. SCR for a census varied around the value of 0.7 without any correlation with air temperature (18-24 oC) and the amount of cloud (0-100%). In order to obtain the optimal interval of the census, we plotted the SCR of one, two, three and four censuses of the same month against the 'number of the censuses per month' (NCM), obtaining a saturating type curve. On the other hand, the cost of the monitoring will increase linearly as NCM increases. If we assume that the economical value of the monitoring scheme for the conservational management of the ecosystem is directly proportionate to the SCR, then the (economically) optimal NCM is obtained as the NCM at which the value (along the ordinate) of the saturating formed curve substituted by that of the (linear) cost curve is at maximum. The optimal NCM (= reciprocal of the optimal interval) changed owing to the way of setting the level of conservational management and to that of the rate of cost.
Climate impacts on a North American dragonfly: evolutionary vs. ecological responses Matthews, John H.
University of Texas, Section of Integrative Biology, Austin, Texas 78712, USA. <johoma@mail.utexas.edu>
Increasing numbers of studies describe climate change impacts on long-distance migration phenology, particularly in vertebrates. Insect migrants are much less well described, and few mechanisms influencing the timing of migration have been posited for any species. Anax junius is a North American dragonfly species with two sympatric dispersal phenotypes. The migrant phenotype appears to engage in long-distance seasonal mass migration, while the resident phenotype remains much closer to its natal pond as an adult. Emergence data collected in southern Ontario in 196869 by Trottier described distinct development phenologies for each phenotype, particularly with respect to the timing of emergence. Mean temperatures and the timing and amount of precipitation have shifted for this population since 1970. By 200305, this populations overwintering resident larvae were emerging later and resident larvae were emerging earlier. Are these changes the result
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of sustained natural selection, or has phenology simply tracked climate variables? Possible influences and mechanisms are considered, including shifts in developmental processes, ambient air versus water temperatures, the dispersal phenology of migrating adults, and regional variation in emergence behavior.
Flying Colours: Five years of research on Odonata in tropical eastern Africa Dijkstra, Klaas-Douwe "KD" B.
Gortestraat 11, NL-2311 MS Leiden, The Netherlands. <dijkstra@nnm.nl>
Over 850 species of Odonata occur in tropical Africa. Together with Viola Clausnitzer I have been working on an identification manual and checklist for the 470 species occurring in the eastern half, performing numerous taxonomic revisions and extensive fieldwork. Having reviewed the status and nomenclature of almost 500 species, it has become apparent that about 13 % of prevailing species names appear to be synonyms, 5 % of species will see a change of their familiar scientific name and about 13 % of country records cannot be validated. This amount of confusion has many causes, of which a previously unrecognised degree of variability in the species is perhaps the most important one. Only about ten eastern African species are new to science. An impression of eastern Africa's odonate diversity is given with highlights from the fieldwork. The endemics of the Ethiopian highlands were photographed and their habitats documented for the first time. The conservation status of Oreocnemis phoenix, a genus and species endemic to Malawis Mount Mulanje, was assessed. Its phenology and ecology is discussed in relation to the mountains climate. The new corduliid Idomacromia jillianae was discovered in Ugandas Impenetrable Forest. A visit to the central Congo Basin in the Democratic Republic of Congo revealed many species not previously found so far east, as well as species new to science. The mighty Congo River itself has an interesting fauna rich in gomphids, which proved rather difficult to study. I conclude with some perspectives for Afrotropical dragonfly research. Priorities lie in the discovery of larvae and the reconstruction of phylogenies. The latter may help to understand the history of the continent and its odonates in relation to its changeable climate, and the mechanisms of species radiations, such as that of the Chlorocyphidae.
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A preliminary phylogenetic hypothesis of Odonata, based on multiple molecular and morphological data sets Kjer, Karl M.; Carle, Frank L. & May, Michael L.
Departments of Ecology & Evolution and Entomology, Rutgers University, New Brunswick, New Jersey 08901, U.S.A. <mimay@rci.rutgers.edu>
Phylogenetic relationships among Odonata at every level from suborder to genus have long been controversial. Recently several authors have applied cladistic methodology to morphological or molecular data in an attempt to resolve some of these uncertainties, but disagreement remains. We here report an analysis based on three separate published morphological data sets in addition to sequence data from nuclear ribosomal DNA and a nuclear protein coding gene from our own work and mitochondrial ribosomal DNA from the literature. Relationships based on these independent data show a high degree of congruence at the family level, although several areas of discordant topology have yet to be resolved. Our results suggest that Zygoptera are monophyletic and are the sister clade to Epiophlebidae + Anisoptera. Lestoidea are basal within Zygoptera, with Hemiphlebia most likely a basal lestoid. A more-or-less traditional Coenagrionoidea is sister to a clade comprising, in approximately basal to terminal order, Megapodagionidae, Pseudolestidae, Amphipterygidae, Diphlebiidae, and Calopterygoidea (although not all these taxa are resolved as monophyletic). Aeshnoidea (Austropetaliidae + Aeshnidae) are basal within Anisoptera. Gomphoidea (= Gomphidae, or possibly Gomphidae + Petaluridae) is then the sister clade to the remaining Anisoptera. This large remaining clade includes (probably) a basal Petaluridae, followed in a pectinate fashion by a paraphyletic "Cordulegastroidea", then Synthemistidae + Gomphomcromiidae, a paraphyletic "Corduliidae", including Macromiidae and a number of genera of still uncertain placement, in addition to Corduliidae s. s., and finally Libellulidae. These results suggest that petiolate wings are plesiomorphic in Odonata and that multiple antenodal crossveins in the calopterygoid zygopteran clade are convergently similar to Anisoptera (or to basal Odonata). Multiple similarities in aeshnoid and libelluloid eyes, male epiproct, planate veins and anal loop, and abdominal carinae are convergent, as are similarities in secondary genitalia and loss of ovipositor in gomphids and libelluloids. The monophyly of Cavilabiata is supported, implying that the sperm pump of higher libelluloids evolved from cordulegastrid-like ancestors. Although it is clear that extensive additional data are needed, we consider most of the basic framework outlined here to be reliable.
Ecomorphology of legs in larval and adult Anisoptera Leipelt, Klaus Guido

Institut fr Geokologie, Technische Universitt Braunschweig, Langer Kamp 19c, D-38106 Braunschweig, Germany. <k.leipelt@tu-bs.de>
In Odonata the transition from ultimate stadium larvae to adults is characterised by ontogenetic changes in morphology, function, behaviour, and habitat. The functions of larval legs vary considerably due to microhabitat occupancy. Therefore, larval Anisoptera can be divided into several larval types according to Corbet (1999, Dragonflies: behaviour and ecology of Odonata; Harley Books, Colchester): claspers climb among the vegetation,
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sprawlers use their long, laterally extended legs to support the body on or among detritus or macrophytes, and burrowers bury themselves in sediment by executing digging movements with the legs. The functions of legs of the adults seem to be similar among Anisoptera; however, two behavioural types can be classified: fliers adopt a vertical posture when resting with the body hanging down, whereas perchers during daylight rest on a perch adopting a horizontal posture with the body supported on the legs. The lengths of leg segments are probably subject to both larval and adult constraints. Therefore, it was hypothesised that ontogenetic changes in lengths of leg segments should occur, which vary between behavioural types. Different larval and adult types should show differences with regard to morphometric leg parameters (the lengths of front, middle, and hind femora, tibiae, and tarsi). Ten species were analysed in this study: three claspers, three sprawlers, and four burrowers; six fliers and four perchers. Morphometric leg parameters were measured in ultimate stadium larvae and adults out of five families (Aeshnidae, Gomphidae, Cordulegastridae, Corduliidae, and Libellulidae; with two species each). Cluster analysis revealed the existence of three different groups with regard to ontogenetic changes of leg segments, one group consisting of the three clasper species (two fliers, one percher), another group consisting of two burrower species (two perchers), and the largest group consisting of two burrowers and the three sprawlers (four fliers, one percher). For instance, the femur/tibia length ratio changed significantly from larvae to adults, the most considerable changes were measured in two gomphid burrower species: in the front and middle legs which are the legs used for digging the femur/tibia ratios changed from in larvae to approximately 1 in adults. Consequently, the larval femur/tibia ratios of the gomphid species were the smallest of all ten species, the ratios of the adults were among the largest.
Critical and consequent taxonomy in Odonata: the European perspective Dijkstra, Klaas-Douwe "KD" B.
Gortestraat 11, NL-2311 MS Leiden, The Netherlands. <dijkstra@nnm.nl>
Worldwide numerous taxonomic problems exist in Odonata, particularly concerning generic placement and the value of subspecies. The problems are caused in part by uncritical and inconsequent taxonomic practice. The seemingly haphazard outcomes have given rise to much resentment among both taxonomists and non-taxonomists for namechanges. Although European Odonata are often considered as well resolved, this is only partly true. With the preparation of a new fieldguide by me and illustrator Richard Lewington for 160 species in Europe, NW Africa and W Turkey, it is timely to review the problems and draw consistent conclusions. Criteria are formulated to assess the value of genera and subspecies. Their suitability is tested on current knowledge of problems like Aeshna isosceles and Cercion lindenii. Especially the merits of small genera within large complexes (e.g. Hemianax in Anax, Chalcolestes in Lestes) seem debatable. New data coming from phylogenetic analysis of DNA should solve most problems in the near future, but more changes are possible, for instance in Aeshna and Libellula. The considerations are enlivened with Lewingtons artwork for the new book.
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Aquatic egg-parasitoids (Hymenoptera) of dragonflies and other arthropods: unique life and flight under water Fursov, Viktor
Institute of Zoology of National Ukrainian Academy of Sciences, iev, Ukraine. <v_fursov@yahoo.com>
The original data on biology and behavior of aquatic parasitic wasps (Hymenoptera), being egg, larval and pupal parasitoids of insects and some other arthropods (Arachnida), living in aquatic and semi-aquatic habitats, are discussed. Original methods of field collecting and rearing of aquatic wasps from eggs and larvae of aquatic hosts are described for the first time. Biology of aquatic wasps is discussed. Aquatic wasps, parasitizing eggs of aquatic and semiaquatic hosts (insects and other arthropods), have such trophic relations with their hosts: Trichogrammatidae, Mymaridae, Eulophidae egg-parasitoids of predaceous diving beetles (Coleoptera: Dytiscidae), beetles-hydrophilids (Hydrophilidae), aquatic bugs (Heteroptera: Nepidae, Naucoridae, Notonectidae), water flies (Diptera: Tabanidae, Sciomyzidae) and dragonflies (Odonata: Aeshnidae, Lestidae, Coenagrionidae, Platycnemididae, Epiophlebiidae); Encyrtidae and Trichogrammatidae egg-parasitoids of sialids (Megaloptera: Corydalidae, Sialidae); Scelionidae egg-parasitoids of water scaters (Heteroptera: Gerridae), dragonflies (Aeshnidae), water flies (Tabanidae), water bugs (Nepidae) and semi-aquatic spiders (Arachnida: Tetragnathidae). It was recorded that aquatic wasps can live completely under the water up to 15 days (Prestwichia aquatica) but they die after 15 minutes being out of the water. Aquatic wasps have unique ability to dive and swim under the water by means of their legs (Prestwichia aquatica, . solitaria, P. multiciliata, Gyrocampa stagnalis, Tiphodytes gerriphagus, Agriotypus armatus). Several species of parasitoids can swim under the water using their wings as the oars (Hydrophylita aquivolans, araphractus cinctus, Aprostocetus natans and Lathromeroidea silvarum). The ability to swim under the water by means of wings was firstly described for 2 species of aquatic wasps (Aprostocetus natans and Lathromeroidea silvarum). The ability of Prestwichia multiciliata to swim under the water by means of legs and active flight in the air by wings was recorded at the first time. The unique aquatic behavior and hosts of Baeus japonicum, Lathromeroidea silvarum, Prestwichia multiciliata, Calotelea shimurai, one species of Oligosita and five species of Aprostocetus were described at the first time. Twenty species of aquatic egg-parasitoids were collected in Ukraine and Russia. Twelve species of aquatic egg-parasitoids (Hymenoptera) were collected by the author in Japan: 2 species of Trichogrammatidae, 2 species of Mymaridae, 2 species of Scelionidae and 6 species of Eulophidae. Six species of aquatic parasitoids were collected in Mexico. Material of parasitoids for present study and observations was collected by the author in Ukraine, Russia, Japan, Mexico, Ireland, England, Hungary.
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On some paintings of Odonata from the late Middle Ages (14th and 15th centuries) Carvalho, Alcimar do Lago
Departamento de Entomologia, Museu Nacional, Universidade Federal do Rio de Janeiro; Caixa Postal 68044, BR 21944-970, Rio de Janeiro, RJ, Brasil. <alagoc@acd.ufrj.br>
European works of art from the late Middle Ages (14th and 15th centuries) displaying insects are rare. The insects more commonly represented are butterflies and flies. Their representations are not merely decorative, having often a symbolic moral meaning in the depicted scene. In general, in this period, butterflies were symbols for the soul and eternal life, and flies were symbols for the brevity of the earthly life. Up to now representations of Odonata of only five different sources of the period were registered. For this essay representations found in six other masterpieces, including paintings and illuminations on books, were considered. The possible symbolic meanings assumed by dragonflies and damselflies during the European medieval period were studied for the first time. Representations of Odonata included in the following masterpieces were described and discussed in this study: Belleville Breviary, Paris (J. Pucelle, ca. 1323-1326); Allegory of good government, Siena (A. Lorenzetti, ca. 1338-1340); The book of hunting of Gaston Phbus, Paris (Anonymous, beginning of the 15th Century); The two lovers, Southern Germany (anonymous, ca. 1470); Hastings Hours, Flanders (anonymous, ca. 1480); Hours of Engelbert of Nassau, Belgium (Master of Mary of Burgundy, ca. 1485). As we might expect, the known representations of Odonata from the late Middle Ages, made in different places and ages, show distinct representational patterns, functions and possible symbolisms. Their inclusion on a work of art can always be interpreted as a symbol to complement or reinforce a painters idea. Thus, their meaning in each of the representations studied seems to be somewhat different. Be it as it may, dragonflies and damselflies probably were considered at that time as messengers of good fortune and signatures of the soul (life after death, change, rebirth), as the butterflies have been since classical antiquity. Nowadays in the western society it is still very common to correlate dragonflies with disincarnate souls. In some of the paintings studied the presence of dragonflies can announce the spring. Captured individuals were probably used as models for some accurate paintings, allowing the identification of some of the genera and species portrayed, as in the case of the painting The Two Lovers, in which a male of Gomphus vulgatissimus can be easily distinguishable. The damselfly painted in the Belleville Breviary (ca. 1323-1326), probably based on a male Calopteryx xanthostoma, is the oldest known European representation of a member of the order Odonata.
Sponsored in part by FAPERJ and FUJB.
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Impact of dragonflies on population suppression of paddy pests in agroecosystem of Kolhapur district, India. Sathe, T.V.; Mundale, Mandar; Bhosale, Y.A. & Margaj, G.S.
Division of Entomology, Zoology Department, Shivaji University, Kolhapur 416 004, India. <pmb_bhoje@yahoo.co.in> Dept. of Agrochemicals and Pest Management, S.U., Kolhapur, India. S.M.B.S., ASC College, Miraj, Dist., Kolhapur, India.
Kolhapur district is the most advanced district of India in agriculture. It has very high rainfall (700-5000 mm). Therefore, paddy and sugarcane are widely cultivated in this district. Biocontrol potential of dragonflies (Order Odonata) have been studied against paddy pests in the fields of Kolhapur. In all 16 species of dragonflies have been reported feeding on rice pests, Lepidopterous borers, leaf roller, and jassids including brown plant hopper (BPH). The important dragonfly species recorded in the paddy fields refers to Diplacodes trivialis (Rambur), D. nebulosa (Fabricius), Neurothermis tullia tullia (Drury), Crocothemis servilia (Drury), Brachythemis contaminata (Fabr.), Orthetrum sabina sabina (Drury), O. chrysis Selys, Acisoma p. panorpoides Rambur, Trithemis aurora (Burmeister), Pantala flavescens (Fabricius), Rhyothemis v. variegata (Linnaeus), etc. Out of which C. servilia and P. flavescens were most dominant species in the rice agroecosystem of Kolhapur district. The above two species significantly reduced the population of BPH and Lepidopteran borers of paddy fields.
On the genus Dubitogomphus Fraser, 1940, their true status and characters Karube, Haruki; Katatani, Naoharu & Kitagawa, Kazuma
Kanagawa Prefectural Museum of Natural History, 499, Iryuda, Odawara, 250-0031, Japan. <qtmfc457@ybb.ne.jp>
We found interesting median sized Gomphidae in the northeast mountain area of Laos in the spring of 2003. They are characterized by having a cross vein in the triangles of hind wings, expanded abdominal ends, ventrally expanded posterior hamuli, and female with a pair of horns on the head. After careful study, it is clear that the species resemble to Davidius? bicornutus Selys, 1878. But, Chao (1990) already has questionned that D.? bicornutus belongs to the genus Davidius. The reason was a rather different venation and peculiar penile organs. Finally, we notice that D.? bicornutus, in fact, should belong to the genus Dubitogomphus Fraser, 1940. Fraser established this new genus for the Indian species Leptogomphus bidentatus Fraser, 1930 in his study of Gomphidae penis. The genus has been known monotypic long time, but, we will move D.? bicornutus to this genus. Here, we report the peculiar characters of Dubitogomphus and also discuss its relationship with other genera.
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Comparative studies on the genital and sub-genital abdominal segments of five species of dragonflies (Anisoptera: Odonata). Thomas, Manu; Gunasekaran & Mohan, Daniel
Department of Zoology, Madras Christian College, Tambaram, Chennai.600 059, India. <mtmathai@hotmail.com>
In the five species of anisopterans, Pantala flavescens (Fabricius), Tholymis tillarga (Fabricius), Brachythemis contaminata (Fabricius), Bradinopyga geminata (Rambur), Diplacodes trivialis (Rambur), the accessory copulatory organs, which situated at the posterior end of the abdomen, are well developed particularly in the case of males. The epiproct, which represent the extension of the eleventh abdominal segment is comparatively larger in males than in females. Compare to the size of the epiproct and paramere, the hypoproct is smaller in all the five species. With respect to hypoproct they are larger in males than in females. The gonopodes of the males of all the five species are well developed exhibiting a pear shaped structure except in Diplacodes trivialis, where it appears as a bean shaped structure. The size of the paramere did not express any uniformity with respect to the sex or size of the species. In Brachythemis contaminata, Bradinopyga geminata, Diplacodes trivialis, the size of the paramere of the females comparatively smaller than in the males, but in Tholymis tillarga the females have a comparatively larger paramere. It is interesting to note that the ninth abdominal segment where the genital opening is situated a pair of small protuberance is observed in the present study.
Allochthonous organic matter as a food resource for aquatic invertebrates in forested streams Graa, Manuel A. S.
Departamento de Zoologia, Universidade de Coimbra, 3004-517 Coimbra, Portugal. <mgraca@ci.uc.pt>
This paper summarises the role of organic matter in the ecology of forested low order streams. Forests are among the most productive systems on Earth. More than 90% of forest primary production will end in detrital pathways, in soil and water. The amount of energy in the form of plant litter entering forested low order streams is several times higher than the energy synthesized by aquatic producers; therefore leaves produced in the riparian zones are a main energy source and decomposition is an important ecological process in those systems. Decomposition is mainly a biological process initiated by aquatic fungi and shredding invertebrates. Those organisms promote the transformation of leaves into fine particles used by bacteria, collectors and filter-feeding invertebrates. Therefore, much of the energy allocated into secondary production is streams has an allochthonous origin. Nutrients liberated as a result of decomposition are used further downstream, in lakes or estuaries by primary producers. The rate at which leaf litter is decomposed is controlled by intrinsic leaf properties (nutrient content, plant chemical and physical defences) as well as environmental factors (e.g. nutrients in water). Perturbations to riparian zones and eutrophication therefore affect decomposition and for this reason, changes in decomposition rates can be used as a functional parameter to assess stream health. Given that the standing stock of leaf litter has a positive effect on leaf consumers, allowing high
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biomass and diversity, it is likely to also affect top predators including odonates; however, the literature on this subject is still scarce.
Abstracts Book 4th WDA International Symposium of Odonatology, Pontevedra 26-30 July 2005 Poster presentations
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POSTER PRESENTATIONS 1. Preliminary observations of reproductive behavior in Arigomphus villosipes (Anisoptera: Gomphidae) McMillan, Victoria E.
Department of Biology, Colgate University, 13 Oak Drive, Hamilton, NY 13346, USA <vmcmillan@mail.colgate.edu>
I studied the behavior of the Unicorn Clubtail (Arigomphus villosipes) at a small artificial pond in New York State from 6 June-8 July 2002. During the day, males adopted perch sites usually 3 m or farther apart on the ground or on rocks along the shoreline. The vast majority of their time was spent perching, interrupted by occasional patrols over shallow water. Perched males also attempted to clasp females appearing at the water. Most reproductive activity occurred between 0900-1630 h. During periods of peak activity, in late morning and early afternoon, 16-20 males were distributed along the shoreline. Perched males occasionally chased other male A. villosipes, as well as other odonates; however, sometimes two males perched close to each other without apparent interaction. Marking records for 33 males showed that most (79%) returned to the pond on one or more subsequent days. Males displayed only weak attachment to perch sites, often occupying 2 or more different areas along the shoreline over several hours on a given day. Female visits to the pond were brief and infrequent; hence, male-female encounters were rare. Females oviposited by flying slowly over shallow water, dipping the abdomen once or twice at multiple sites. Mate-guarding was never observed. Females that were ovipositing near perched males were chased and sometimes taken into tandem, whereupon the pair left the pond. Copulation, which probably occurred in the surrounding field, was not observed. Some females escaped male harassment by ovipositing at times when no males were present, or in locations 3 m or farther from perched males. Reproductive behavior of A. villosipes is discussed with reference to mating systems in other gomphids.
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2. Site fidelity, mating success and reproductive strategies in males of Libellula fulva (Odonata: Libellulidae) Nagy, H. Beta1; Lszl, Zoltn2; Szllassy, Nomi3; Szkely, Annamria4 & Dvai, Gyrgy1
1 Dept. of Hydrobiology, University of Debrecen, 4032 Debrecen, Egyetem tr 1, Hungary. <nagy.beata@gmail.com> 2 Dept. of Ecology, University of Debrecen, 4032 Debrecen, Egyetem tr 1, Hungary 3 Faculty of Psychology and Science of Education, Babe-Bolyai University, RO-400015 Cluj, Str. G. Bilacu 24, Romania 4 Faculty of Biology and Geology, Babe-Bolyai University, RO-400015 Cluj, Str. G. Bilacu 44, Romania
During two seasons (2002-2003) we studied a closed Libellula fulva population along a small creek in Eastern Hungary. The territorial behaviour of males was observed with binoculars. Studied individuals were marked on their right wings. We marked 169 males in 2002, and 186 males in 2003. Movement of marked males was recorded along a 350 meter natural section of the stream that was marked every five meters. We investigated the site fidelity of the males by the localisation (LI) and the site fidelity (SFI) indices. In 2002 we calculated the indices for 26, in 2003 of 46 males. We observed that territorial males defended not just one, but simultaneously two or three territories. The aim of the study was to determine which group of territorial males (with one, two or three defended territories) insists more on their territory (has higher habitat fidelity). We also studied the differences in the mating success of territorial and non-territorial males. Further objective was to explore the differences of territorial and non-territorial males in reproductive strategies. We found that SFI was the highest in the group of males, which defended three territories, and LI was highest for males, which defended only one territory. Mating success of territorial males was significantly higher than the mating success of non-territorial ones. Territorial males showed satellite behaviour more frequently than non-territorial males. It seems that the satellite behaviour is a successful alternative reproductive strategy of both territorial and non-territorial males in the species Libellula fulva.
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3. Mating behaviour in damselfly Ischnura senegalensis (Rambur) Sathe, T.V.1; Bhoje, P.M.2, Kavane, P.3, Shinde, Kiran4; Yadav, R. P.4 & Pandharbale, A. R.1
1 Entomology Division, Department of Zoology, Shivaji University, Kolhapur 416 004, India. <pmb_bhoje@yahoo.co.in> 2 Y.C. College, Warananagar, India. 3 Vivekanand College, Kolhapur, India. 4 DKAS College, Ichalkaranji, Dist, Kolhapur, India.
The damselfly Ischnura senegalensis (Rambur) (Odonata: Agrionidae) is an important biocontrol agent of Lepidopteran pests and jassids of rice in agroecosystem of Kolhapur, India. Ethology of a pest biocontrol agent has a direct influence on the population of pest species and the success of the biocontrol programme. Therefore, mating behaviour in the above species has been studied. Males are blue green with black lines and females are reddish brown with dark lines on the thorax. The pair hold their wings against each other while resting. The mating behaviour consists of a typical chain of event viz. territorial behaviour, courtship and recognition, grasping and clasping, tandem, intramale sperm translocation, copulation invitation, copulation wheel position, tandem after copulation and separation. Details of this mating behavioural chain are discussed in this paper.
4. Egg distribution, mate-guarding intensity and offspring conditions in dragonflies Schenk, Kamilla
Institut fr Geokologie, Technische Universitt Braunschweig, Langer Kamp 19c; D-38102 Braunschweig, Germany. <k.schenk@tu-bs.de>
How is egg size, larval size, and egg development time related to oviposition site selection and mate guarding in dragonflies? Behaviour studies showed species that oviposited mainly in tandem flight spread their eggs over several ponds (spatial risk spreading). Other species performed non-contact guarding and localised their eggs usually in a single pond only (distinct habitat selection). The longer females oviposited at one place the higher the risk caused by disruption by males or increasing predation. Therefore, I proposed that in species layed their eggs to a distinct oviposition habitat the eggs of a clutch laid first should be the largest, which I interpret as indicator for high condition. Species that perform risk spreading should distribute large eggs randomly over all oviposition sites. The hypothesis was tested and confirmed at artificial ponds in Namibia and Germany using experimental manipulations of oviposition.
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5. Fat reserves and wing spot size, but not wing spot and body colour intensity, are related to male success during territorial contests in Hetaerina americana Contreras-Garduo, Jorge; Buzatto, Bruno Najera, Karla & Crdoba-Aguilar, Alex
Previous work has shown that wing pigmentation spots in males of the damselfly H. americana, are sexually selected: territorial males have larger wing spots than nonterritorial males. This is possibly related to the fact that pigmentation may be an indicator of fat reserves. In this work we have investigated the role of spot size, spot and body colour intensity and fat reserves in determining male contest success in H. americana. We determined male territorial success in 15 natural fights in Tehuixtla, Mxico, in April and May, 2005 and related this to spot size, spot and body colour intensity and fat reserves of both opponents. We recorded colour intensity and hue using live animals immediately after the encounter using a field spectrophotometer. Fat reserves and spot size were determined in the laboratory using standard damselfly methodologies. Our results indicated that only fat reserves and spot size, but not wing spot and body colour, were related to successful males after an encounter. This suggests that male assessment of opponents is related to wing spot size as a possible indicator of fat reserves. That colour intensity is surprising given recent findings in other damselfly species and the fact that this variable seems to vary considerably during adulthood.
6. The effect of thermal pollution on dragonfly populations Mihokovic, Nino1 & Slavikovski, Ana2
1Bribirska 2Vida
39, HR-10000 Zagreb, Hungary. <nino.mihokovic@inet.hr> Dosena 41, HR-10000 Zagreb, Hungary.
The effects of thermal pollution of Zagreb citys area are correlated with dragonfly moulting time, population age and composition. A location under the influence of thermo electric power plant, urban and industrial environment is compared to a location with similar population composition isolated from allochthonous heat sources. Five stations on each location are compared between each other using Sorenson similarity coefficient. Capture Mark Recapture methods are employed in order to aid in population ageing.
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7. Evaluation of the odonata community in Galician rivers (NW Spain) that are affected by hydroelectric power stations Rodrguez-Guntn, I.; Prez-Bilbao, A.; Gonzlez, A.; Alonso, A. & Garrido, J.
Departamento de Ecologa y Biologa Animal, Facultad de Biologa. Universidad de Vigo, Lagoas Marcosende, 36200 Vigo, Pontevedra, Spain. <iriaguntin@uvigo.es>
Dragonflies are a well studied group of insects in the Iberian Peninsula. In Galicia their knowledge has increased during the last few decades and nowadays its estimated that 44 of the 75 species that are identified in the peninsula, have been recorded in this autonomous region. The results of a study done in Galician rivers that have hydroelectric power stations are presented in this paper. The objective was to evaluate the possible effects of these constructions on the macroinvertebrate communities, and especially in our case, on the Odonata species. The study was done from 1998 to 2002 in five Galician rivers (Deva and Tuo in Ourense; Tambre in A Corua; and Deva and Tea in Pontevedra). We planned seasonal samplings during a year for each river, and there were established different sampling points according to the number of power stations that each river has. To follow a pattern, the sampling points were always located upstream the station, immediately after it and a few kilometres downstream. 460 odonata larvae, that belong to 3 families of the Zygoptera suborder and the other 3 to the Anisoptera suborder, were captured. The identified species were the following ones: Calopteryx virgo (L. 1758), Calopteryx haemorrhoidalis (Van der Linden, 1825), Calopteryx splendens Harris 1782, Lestes viridis (V. L. 1825), Platycnemis sp. (Burm.), Cordulegaster boltonii (Donovan 1807), Boyeria irene (Fonsc. 1838), Gomphus simillimus Selys, 1840, and Onychogomphus uncatus (Charp., 1840). The abundance and richness were evaluated, and also the spatial variation of the odonata communities.
8. Comparative molecular genetic analysis of the population structures and dynamics of two aeshnid species (Odonata: Aeshnidae) in Namibia Timm, Janne & Hadrys, Heike
Inst.fr Zellbiologie und Tierkologie, Tierrztliche Hochschule Hannover, Bnteweg 17d, 30559 Hannover, Germany. <janneket@gmx.de>
Molecular genetic methods have become an important tool for the measurement of biological diversity on the population and species level. To evaluate and conserve biodiversity it is necessary to understand about genetic diversity and population structures of species. In this work, the genetic diversity and population structures of two dragonfly species, Anax imperator and Anax speratus (Fam.Aeshnidae), were studied, which show different degrees of habitat specialization and dispersal potential. The geographical centre of the study was the arid country Namibia, where a high diversity of odonates is found despite of its dry climate. For the population genetic analyses a fragment of the mitochondrial ND1-Region was sequenced, genealogical networks constructed and the variability and structuring statistically calculated. As results the habitat specialist Anax speratus shows very small genetic variability but a strong differentiation between two populations that indicates a restricted gene flow. Because of this small variability and the isolation of the populations it is likely that this species is very sensible to changes in their
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habitat. The generalist Anax imperator on the contrary shows a high genetic variability and no significant structuring of the populations because of its high dispersal potential. To conclude, this study shows the influence of the degree of habitat specialization on the genetic diversity and the population structures in these two aeshnid species.
We acknowledge the support of the BMBF (BIOTA-Project S08).
9. Hybridisation and inheritance of female-limited colour polymorphism in two ischnurid damselfly species (Odonata) Van Gossum, Hans1, Snchez-Guilln, Rosa Ana2 & Cordero Rivera, Adolfo2
University of Antwerp, Evolutionary Biology Group, Groenenborgerlaan 171, B-2020 Antwerp, Belgium. <Hans.VanGossum@ua.ac.be> 2 Grupo de Ecoloxa Evolutiva e da Conservacin, Departamento de Ecoloxa e Bioloxa Animal. Universidade de Vigo. EUET Forestal, 36005 Pontevedra, Galiza, Spain.
1
Female-limited colour polymorphism is frequent in many species of Odonata. Ischnura elegans exhibits three colour morphs, one male-like coloured (andromorph) and two gynomorph brown morphs (infuscans and rufescens-obsoleta). A total of 19 progenies obtained from once-mated females were reared in the laboratory in three generations. Results indicate that the colour morphs are controlled by the same genetic system as previously described for I. graellsii, i.e. an autosomal locus with female-limited expression and with three alleles with a hierarchy of dominance (pa>pi>po). Five interspecific crossings between female I. graellsii and male I. elegans, five crossings between hybrid females and male I. elegans and one crossing between female I. graellsii and a hybrid male further confirmed that the genetic system is the same in both species. A survey of morph frequencies in NW Spain revealed that I. elegans shows high variability in andromorph frequency (4-91%) between near populations, whereas in I. graellsii andromorphs never are the majority morph (5-40%). The highest andromorph frequency in I. graellsii was found in populations closest to a locality where both species have hybridised, and that now has the highest andromorph frequency of I. elegans. We hypothesise that I. elegans genes have been incorporated into the genome of I. graellsii and increased andromorph frequency in the latter species. Low andromorph frequency in I. elegans seems also related to the influence of I. graellsii genes. Therefore we suggest that hybridisation between both taxa is contributing to the temporal maintenance of contrasting andromorph frequencies in nearby populations.
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10. Genetic consequences of a man-made bottleneck in Orthetrum coerulescens: A microsatellite system to study fine scale population dynamics Wargel, Antonia, Giere, Sandra & Hadrys, Heike
ITZ, Ecology and Evolution, TiHo Hannover, D-30559 Hannover, Germany. <A.Wargel@gmx.de>
Irrigation ditches are often the only viable breeding sites left for riverine dragonfly species in developed landscapes. Dredging of irrigation ditches is a common anthropogenic deterioration of invertebrate communities. We developed a microsatellite system for the keeled skimmer (Orthetrum coerulescens) to investigate whether dredging of its breeding habitat leads to immediate and long-term genetic traces in the population structure. We combined monitoring data on population size with genetic data on population structure before and after dredging of the breeding site (a small irrigation ditch in southern France). Fine scale population studies to monitor a direct genetic response to environmental changes are mostly limited by a low resolution of genetic markers. The high sensitivity of microsatellite markers allows fine scale estimates of local dispersal patterns. For O. coerulescens, typing of 7 microsatellite loci resulted in an overall number of 81 alleles. The results show that despite the fast recovery in terms of population size and reproductive output, the population was affected by a cryptic loss of genetic diversity. The loss of 10 private alleles and a mode shift in allele frequencies demonstrates the impact of dredging on O. coerulescens.
11. The effect of larval density on female polychromatism and body size in Ischnura graellsii Gonzlez de Castro, Ins1; Van Gossum, Hans2 & Cordero Rivera, Adolfo1
1 Grupo de Ecoloxa Evolutiva e da Conservacin, Departamento de Ecoloxa e Bioloxa Animal, EUET Forestal, Universidade de Vigo, Campus Universitario, 36005 Pontevedra, Spain. 2 University of Antwerp, Evolutionary Biology Group, Groenenborgerlaan 171, B-2020 Antwerp, Belgium.
In Ischnura graellsii, (Odonata: Coenagrionidae), three morphs can be identified: the androchrome morph, that is male-like coloured; and two gynochrome morphs (aurantiaca and infuscans), with brown coloration, completely different from males. In this paper we examine if the colour morph has an impact on the development and competitive ability of larvae. We obtained eggs from from adults collected in the field, and incubated them in the laboratory. Larvae were randomly distributed in three groups with different density (300, 600 and 1200 at the start of the experiment), and four replications (aquaria) per treatment, and the same amount of food (40 ml/day of Artemia and some aquatic oligoqueta in the last stages) was distributed among the three groups. Upon emergence, adults were measured, sexed and the colour morph of females recorded. A regression analysis indicates that treatment had a significant effect on male length, since it decreases when population density increases. The treatment has also an effect on the sex ratio, increasing male proportion at high density. There was no significant effect of treatment on androchrome frequency. These results suggest that there are little or no pleitropic effects of the colour gene on larvae competitive ability
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12. Preliminary results of a multi-year study of phenology and development of Anax junius in Maryland, U.S.A. May, Michael L.
Department of Entomology, Rutgers University, New Brunswick, New Jersey, 08901, U.S.A. <mimay@rci.rutgers.edu>
In the northeastern United States, mass migrations of the common green darner dragonfly, Anax junius, are regularly observed during the autumn, and recent observations (Orr, 1996; Russell, et al., 1998) confirm that a return migration occurs in spring, although movements are less obvious. In southern Canada, at least, migrant and resident populations of A. junius apparently were found to be quite distinct and possibly reproductively isolated (Trottier, 1966, 1971). Observations of adult activity and emergence at Patuxent National Research Refuge in Maryland, U.S.A. (30.02 N, 76.47 W) suggested the same might be true there (Orr, 1996, and pers. comm.). To test Orr's conclusions, I have collected larvae at two ponds on the refuge, using a standard aquatic dipnet, once or twice each month from April to October or November during 2001 and 2003-2005). Head width, wing pad length, and total body length are measured in the field, then most of the larvae collected, 10-30 for each sample, are returned to the ponds. Plots of these morphometric data over time throughout the adult flight season indicate the existence of identifiable migratory and resident cohorts. However, emergence data indicate that adult flight seasons overlap extensively, thus permitting gene flow between them. This is in accord with earlier findings that putative migratory and resident individuals do not differe consistently in genetic characteristics (Freeland, et al., 2003). Also of interest is the obserevation that the extent of year-to-year variation in population size and relative success of overwintering (resident) vs migrant Anax junius is much greater than has been appreciated heretofore.
13. Relative frequency of Ischnura elegans and I. graellsii (Odonata: Coenagrionidae) in the Galician coast Snchez-Guilln, Rosa Ana & Cordero Rivera, Adolfo
Grupo de Ecoloxa Evolutiva e da Conservacin, Departamento de Ecoloxa e Bioloxa Animal, Universidade de Vigo, EUET Forestal, 36005 Pontevedra, Spain. <rguillen@uvigo.es>
When reproductive isolation is incipient, mating between related species can produce viable hybrids that give place to new species, or that allow the absorption of a species by another. We have studied the geographical distribution of I. elegans and I. graellsii in NW Spain over the last years. We found that both species are rarely sympatric, and that some localities that had graellsii populations are now colonized by both species or only have elegans. Under laboratory conditions 90% of matings are between male elegans and female graellsii, and the same unidirectional mating has been observed in the field. Both types of matings produce hybrids with low viability and fertility, that can be backcrossed with the parental species. In this paper we review all available data on the relative frequency of both species in Galician localities. In at least two localities studied during 2003, monthly samples indicate that elegans is increasing very fast, and this suggests that graellsii might disappear from some localities in the next years.
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14. Present dragonflies in As Gndaras de Budio. Site of Community Importance (Nature 2000 Network) Alonso, A.; Garrido, J.; Prez-Bilbao, A.; Gonzlez, A. & Rodrguez-Guntn, I.
Campus As Lagoas. Universidad de Vigo. 36310 Vigo, Spain. <aiap@uvigo.es>
The wetland As Gndaras de Budio is included in the Nature 2000 Network of Galicia. It is one of the 59 open spaces that has been declared as a special place of protection of the natural values by the Order 72/2004 of April the 2nd. It is also included in the list of Sites of Community Importance of the Atlantic biogeographical region (DOCE L387 of 29/12/2004). As Gndaras de Budio are located in the town councils of O Porrio, Salceda de Caselas and Tui (Pontevedra), all in NW Spain. This place is between the coordinates 837 W and 425 N., and has a size of 727 ha and 25 m of altitude. It is a marshy area with an important riparian forest. The aquatic and peat bog vegetation is also well represented. We can find natural eutrophic ponds with Magnopotamion or Hydrocharition vegetation. The Odonata are a very interesting group for the study and conservation of the aquatic systems because of their abundance, diversity, conservation interest or ecological requirements. Their value as good indicators of biodiversity and habitat conservation is well documented (Van Tol & Verdonk, 1988; Collins & Thomas, 1991; Samways, 1994); even the Coenagronidae family has been proposed the best indicator taxa of invertebrate biodiversity in ponds (Briers & Bigss, 2003). The dragonflies are the invertebrate group that comparatively has the highest representation in catalogues and threatened species agreements, with 6 species in the different categories of the National Catalogue. This is one of the few groups of invertebrates that has been the objective of specific actions for their diversity maintenance and increasing, that consists in the recuperation, creation or modification of wetlands, mainly ponds, in protected natural sites (Usher & Jefferson, 1991).
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15. Present knowledge on the Odonata of Serra da Estrela Natural Park (Portugal) Ferreira, Snia & Grosso-Silva, Jos Manuel
CIBIO/UP - Centro de Investigao em Biodiversidade e Recursos Genticos, Universidade do Porto, Campus Agrrio de Vairo, 4485-661 Vairo; Portugal. <hiporame@gmail.com>
The Serra da Estrela Natural Park, a mountainous protected area located in central Portugal, includes a remarkable variety of ecosystems which supports the richest regional biodiversity known in the country, with more than 1000 plant species and over 2300 animal species recorded (more than 2100 of which are invertebrates). The Odonata fauna of the Park is currently being studied through literature and field research and a summary of the results obtained so far is presented, including an historical overview of the study, a list of the species already recorded and distribution maps.
16. Odonata of continental Portugal: mapping the knowledge and identifying geographical gaps Ferreira, Snia1; Grosso-Silva, Jos Manuel1; Soares-Vieira, Patrcia2 & Sousa, Pedro1
1 CIBIO/UP - Centro de Investigao em Biodiversidade e Recursos Genticos, Universidade do Porto, Campus Agrrio de Vairo, 4485-661 Vairo, Portugal. <hiporame@gmail.com> 2 Rua Jos Pedro Ins Canadas, n. 16 - 1. Dto.; 2040-326 Rio Maior, Portugal.
The Portuguese fauna of the order Odonata comprises 65 species, most of which were first recorded from the country during the 19th century. Currently known species distributions rely mostly on local studies, many of which are fairly recent, although a number of more general old studies also exist. In this contribution we analyse the distribution of the recorded species richness on a 50x50 Km UTM grid in order to assess the state of the knowledge on Odonata diversity in a geographical basis and to identify the areas in which further sampling is urgent.
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17. Dragonflies and damselflies of a montane tropical rainforest in Papua New Guinea Oppel, Steffen
Wildlife Conservation Society, PO Box 277, Goroka, EHP, Papua New Guinea. <steffen.oppel@gmx.net>
The dragonfly fauna of Papua New Guinea is species rich, but human population growth and modification of primary rainforests may lead to an unknown loss of species diversity. In this study I sampled odonates in a pristine montane rainforest in Papua New Guinea over several months, and recorded habitat characteristics for all encounters with adult odonates. Using ordination techniques I then classified the odonate fauna into assemblages correlated with environmental factors. I derived 10 different habitat types and then compared the habitat use of two odonate communities in natural and modified forest in order to determine which habitat types are most vulnerable to forest modification. The modified forest community was located in a village, which is representative for widespread forest modification caused by human subsistence gardening. In the pristine rainforest I found 61 species, of which 40% were considered as rare. Cluster analysis identified seven distinct assemblages associated with permanent rivers and creeks, temporary streams, puddles, or permanent standing water. Shading, water speed and water permanence were important factors distinguishing the assemblages. Anisoptera were absent from three habitats in the forest interior with temporary water sources. Species associated with temporary water sources and other microhabitats in the forest interior are presumed to be dependent on the humid conditions of the rainforest. The pristine forest community had a higher diversity and evenness than the village community, and I collected only 37 species in the village. I found 78 species of 13 families in both sites combined, of which 21 were shared between the two areas. Both sites had about three times as many Zygoptera as Anisoptera. Shared species were mostly of the families Libellulidae and Megapodagrionidae, whereas most members of Platycnemidae and Coenagrionidae were restricted to either site. The most important habitats in the village were open sunny areas, artificial ditches, and small permanent creeks, whereas the most important habitats in the pristine forest were running waters of different size as well as the forest interior. Based on the habitat preference of species in the pristine forest, species inhabiting temporary water sources under closed canopy rainforests are most vulnerable to forest modification. They comprised a third of the forest community, and I estimate that at least 25% of pristine rainforest species will disappear following human-induced forest alteration. The preservation of PNG`s highly diverse odonate fauna will depend on the conservation of intact rainforest ecosystems in all parts of the country.
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18. Biodiversity of dragonflies (Odonata) from Western ghats of Maharashtra, India Sathe, T.V.1; Shinde, Kiran1; Bhoje, P.M.2, Thite, H.S.3 & Patil, R.G.4
Division of Entomology, Department of Zoology, Shivaji University, Kolhapur 416 004, India. <pmb_bhoje@yahoo.co.in> 2 Y.C. College, Warananagar. 3 B.P.A.S. College, Angar, Solapur. 4 L.B.S. College, Satara, India.
1
Biodiversity of dragonflies of Western Ghats has been studied in this paper since Western Ghats is among the important global hot spots of biodiversity. Biodiversity protection and conservation is on national and international agenda. Biodiversity is responsible for sustainable development of a region or a country and secondly, dragonflies are potential biocontrol agents of mosquitoes, lepidopteran pests, jassids, etc. In all, 92 species of dragonflies have been reported from Western Ghats of Maharashtra. The important genera includes Lestes, Esme, Disparoneura, Archibasis, Cariagrion, Ischnura, Agriocnemis, Libellago, Macrogomphus, Microgomphus, Anax, Macromia, Orthetrum, Diplacodes, Neurothemis, Trithemis, Pantala, Pseudagrion, Onychargia, Copera, Caconeura, Phylloneura, Prodasineura, Vestalis, Burmagomphus, Asiagomphus and Hemicordulia; out of which Caconeura, Asiagomphus and Macromia genera were represented by rare species while, Phylloneura genus was only confined to Western Ghats and other genera were common in Ghats.
19. On the available data concerning the Odonata of Peneda-Gers National Park (NW Portugal) Soares-Vieira, Patrcia1; Grosso-Silva, Jos Manuel2 & Ferreira, Snia2
Rua Jos Pedro Ins Canadas, n. 16 - 1. Dto.; 2040-326 Rio Maior; Portugal. <pmsvieira@yahoo.com> CIBIO/UP - Centro de Investigao em Biodiversidade e Recursos Genticos, Universidade do Porto, Campus Agrrio de Vairo, 4485-661 Vairo; Portugal.
1 2
Peneda-Gers National Park, which is situated in the North-west of Portugal, bordering Galiza, covers an area of more than 70.000 hectares and includes several mountain ranges (Amarela, Gers, Peneda and Soajo) as well as two high plateaus (Castro Laboreiro and Mourela). The Park is located in the highest rainfall area of continental Portugal, resulting in an intricate network of rivers, streams, and ponds that harbour a rich insect fauna. In this contribution we present data on the Odonata fauna of the Park collected during two invertebrate studies carried out in 2002 and 2003. Before these studies, the Parks recorded Odonata fauna comprised 18 species, to which 5 more were added by the authors. A list of the 23 species and distribution maps of all the species collected during the studies are presented.
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20. A preliminary analysis of odonate species richness in Galiza (NW Spain) Azpilicueta, Mnica; Rey Ra, Carlos; Docampo Barrueco, Francisco; Rey Muiz, Xos Carlos & Cordero Rivera, Adolfo
Grupo de Ecoloxa Evolutiva, Departamento de Ecoloxa e Bioloxa Animal, Universidade de Vigo, E.U.E.T. Forestal, Campus Universitario, 36005 Pontevedra, Galiza, Spain. <adolfo.cordero@uvigo.es>
The analysis of distribution data of odonates in NW Spain indicates the presence of 48 species. Four species (Macromia splendens, Oxygastra curtisii, Gomphus graslini and Coenagrion mercuriale) are protected under the European Habitats Directive and Spanish Law. Localities of collection of specimens collected between 1978 and 2004 were situated in UTM squares of 1010 km to produce a map of species richness for the region. Additionally, we introduced all localities (UTM 11 km) where protected and rare species were found in a GIS, on a map of the Natura 2000 network of the region. Our results indicate that O. curtisii and C. mercuriale are common in NW Spain. We identified as local rare taxa Brachytron pratense, Aeshna affinis and Erythromma viridulum, because they were found in less than 10 squares, and are also relatively rare in the Iberian peninsula. Our analysis reveals that the knowledge of this group is clearly fragmentary, with most records concentrated on the coastal region, and very few squares sampled more than 20 times, the minimum to obtain reliable data. We therefore need a systematic sampling of the region to properly identify areas with high species richness. This will be undertaken by the Galician Natural History Society.
21. Contribution to knowledge of the biology of Onychogomphus costae Slys, 1885 (Odonata: Gomphidae) in southern Spain Cano-Villegas, Francisco J. & Ferreras-Romero, Manuel
Departamento de Ciencias Ambientales (Zoologa), Universidad Pablo de Olavide, Ctra de Utrera km 1, 41013 Sevilla, Spain. <fjcanovi2@hotmail.com>
Onychogomphus costae has a restricted west Mediterranean distribution, occurring only in the Iberian Peninsula (Portugal, Spain) and North Africa (Morocco, Algeria, Tunisia). Its biology has been very little studied. Larvae live in streams and rivers with a moderate current; also occur in slower-flowing, deeper reaches of polluted medium-big rivers. This study was carried out in reaches of the river Guadalquivir which cross the city of Cordova, southern Spain (37 53N, 4 47W, 110 m a.s.l.); its maximum width is ca. 300 m and mean depth bigger than 5 m. The bed of the river is made of boulders covered with fine sedimentary mud. From March to September 2003, 15 visits were effectuated to observe adults; in six of these visits larvae and exuviae were found. In the laboratory, the head width (HW; i.e. the maximum distance between the lateral margins of the compound eyes) of each larva and exuviae, and the length of the metathoracic (hind) wing sheaths (WS), if present, were measured to the nearest 0.1 mm using a Nikon binocular microscope with an eyepiece micrometer. The number of abdominal segments covered by the metathoracic WS was also recorded. On the basis of HW and WS length each larva and exuvia was either assigned to one of the last five stadia or designated a 'smaller larva'. In those samples collected on March 23 and September 24 final stadium larvae (F-0) were found; likewise in that March sample 'smaller larvae' were collected too. Probably there is not egg diapause
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(since during late winter there are F-2 and F-3 larvae, presumably, of the junior cohort), larval growth take place during the autumn, winter and spring, larvae can enter F-0 in late summer (September) and autumn, then final-stadium diapause (short-day diapause) and emerge in their second spring, thus exhibiting a semivoltine life cycle; great part of the population could perform the development as 'spring species' (sensu Corbet 1964). Flight period (33 adults were recorded) from middle May to late August. Mostly adults were seen on dry, yellowish grass, in accordance with the mimetic colours of this species, few times flying over the water of the river.
22. Pond water regime and competition as key factors in the presence and life history of two Lestes damselflies (Odonata: Lestidae) Torralba Burrial, Antonio & Ocharan, Francisco J.
Dpto Biologa de Organismos y Sistemas. Universidad de Oviedo. 33071 Oviedo, Spain. <antoniotb@hotmail.com>
To understand physical and biological factors is necessary for understanding community structure. Lestes virens and Lestes barbarus are damselflies characteristic of temporary ponds, which can coexist or not. In this field study, we analysed pond water regime and damselfly phenology (adult emergence) in ponds from a shire of Huesca (NE Spain). Our results show: (1) a direct effect of pond water regime on the presence of L. barbarus, it not occurring at ponds that dry-up before mid-June because it cannot complete its larval development; and (2) an indirect effect of pond water regime on L. virens phenology: it emerges later in ponds that dry-up later because if coexist with L. barbarus there is competition between the species which delays the life cycle of L. virens.
23. Expansion of Crocothemis erythraea in Ukraine Khrokalo, Lyudmyla & Matushkina, Natalia
Zoology Department, Biology Faculty, Kyiv National Taras Shevchenko University ; Volodymyrska str. 64, Kyiv, Ukraine 01033. <lkhrokalo@mail.ru>
Crocothemis erythraea is a Palaearctic-Afrotropical-Oriental species, which is distributed in Europe, Transcaucasus, the Near East, Mid- and Central Asia, Northern and Tropical Africa, China and Northwestern India. In Europe the Scarlet Dragonfly occurs predominantly in the Mediterranean area. Expansion of this species towards the north, colonization of new biotopes and increase of population size in higher altitudes in Europe have been noticeably recorded during the last 10-15 years. Researchers (Ott, 1996, 2001) consider this phenomenon as consequence of global climatic changed and propose C. erythraea as good indicator for this. In Ukraine the Scarlet Dragonfly was collected in southern and southwestern areas in the beginning and middle of the XXth century (Brauner, 1902, 1903; Artobolevsky, 1029; Bezvali, 1932; St. Quentin, 1933; Grabar, 1933); Pavlyuk (1990) collected this species in 1968 and 1973. During last two decades C. erythraea was recorded in Ukraine in outermost southwest (Gorb, Ermolenko, 1996), in Crimea peninsula (Kiseleva, Vershitsky, 1998; Prokopov, 2003) and, beside that, towards to north
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and east, namely central regions (Barsov, 1987), eastern (Martynov, Martynov, 2003), northern (Tytar, 2003) and northeastern regions (Khrokalo, 2000, 2004). During 2001-2004 we registered new points of distribution of this species in Ukraine (in Crimea, Odessa, Vinnytsya, Cherkasy, Chernihiv and Kyiv administrative regions). Thus, according to original and literature data, we established the fact of expansion of C. erythraea toward north and east of Ukraine during last few years. Our observations revealed that Scarlet Dragonfly showed reproduction and established autochthonous populations in the northern Kyiv region.
24. Sensitivity and economy of monitoring the environment of a large lake using adult dragonflies Kurauchi, Yohei1,2 & Ubukata, Hidenori1
1 2
Hokkaido University of Education, Kushiro, Hokkaido, 085-8580 Japan. <s0520231@ipe.tsukuba.ac.jp> Present address: University of Tsukuba, Tsukuba, Ibaraki, 305-8572 Japan.
In general, larger lakes have higher heterogeneity of environmental conditions. When the environment conditions of such a lake is to be monitored using Odonata, this heterogeneity should be taken into account. It is also necessary to investigate to what extent a monitoring using dragonflies is sensitive to the change of environmental conditions of the lake. Keeping these necessities in mind, we executed intensive censuses of dragonflies at Lake Takkobu (Hokkaido, Japan), which has a contour of about 5 km and maximum depth ca. 2 m and has been suffering eutrophication presumably due to the intensification of stock and dairy farming. We set up 11 quadrats (10 m long, 2 m wide on the water) along the shore, where we censused adult dragonflies six to eight times a month from mid June to early November 2004. Besides, we censused larvae at each quadrat in October, and also exuviae at a quadrat on the shore adjacent to each quadrat for adult sensus. The 'composition of the assemblage of adult dragonfly' (CAA: ratio of the number of individuals of each species to the total species censused throughout the season at each quadrat) considerably differed among the quadrats. The result of a detrended correspondence analysis (DCA) with the CAA of each quadrat showed a two dimensional ordination with the primary axis corresponding to the north-south arrangement of the quadrats at the lake and the secondary, independent axis corresponding to the east-west arrangement of the quadrats. The result of a canonical correspondence analysis (CCA) suggested that the change in the CAA along the north-south axis of the lake strongly correlated with the coverage of the floating-leaved plants. For making the monitoring reflect the CAA of the lake as a whole and the CAA at certain sites representing the heterogeneity, we obtainend the following monitoring scheme based on graphical analyses. The monitoring should be done three times a month at two particular sites at the lake. Thus, this scheme will make the monitoring economical and also practical by involving the heterogeneity of the lake environment. A few dragonfly species were regarded as those which virtually didn't emerge from the lake and flew onto the lake from other habitats. We conclude that the reliability of the monitoring would be enhanced if the larvae and exuviae are censused beside censusing adults.
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25. Flight peak trends for dragonflies from the Netherlands Termaat, Tim1, Ketelaar, Robert1, 2 & de Vries, Henk1
1 Dutch Butterfly Conservation, PO Box 506, NL-6700 AM, Wageningen, The Netherlands. <Tim.Termaat@Vlinderstichting.nl> 2 Present address: Dutch Society for the Conservation of Nature, PO Box 494, NL-5600 AL, Eindhoven, The Netherlands
A flight peak analysis was made for several dragonfly species in the Netherlands. Records from the National Dragonfly Database were used to calculate the flight peak, defined as the mean day on which the species were recorded each year in the period 1980-2004. The results show a considerable shift in flight peak for most examined species, up to an advancement of 18 days. The increase of spring and summer temperatures are an obvious driving factor for these results, although some effect of increased recording intensity is also expected. The results are discussed in relation to the observed expansion and decline of different species. Further research on the relation between climate change and trends in flight peaks can help to predict future scenarios of dragonfly phenology and their consequences.
26. Morphological characteristics of odonate eggs and early instar larvae specific to taxa Watanabe, Yoko
4-14, Nishida-cho, Nishinomiya-shi, Hyogo, 662-0034, Japan.
<mer-yoko.watanabe@nifty.com> There has not been much research conducted into the life cycle of a dragonfly from its genesis through its emergence even though it is quite important. As a result of an examination of the eggs of close to 130 species among the nearly 200 Japanese odonates, it was revealed that they have features specific to taxa. Many of them had a special structure outside their eggshells and the larvae of some odonates were observed to bear morphological characteristics which appeared only in their early instars. Such information obtained from the eggs and early instar larvae has a high potential to contribute to the taxonomy of odonates. It is not yet very easy to speculate on the relationship between such characteristics and the phylogeny or the adaptation to its environment based only on Japanese species. Further accumulation of knowledge in this field is expected.
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27. Interannual change measurement of the Odonata community existing in a Mediterranean river which put up with the by-product of a heavy mining accident happened in April 1998 (river Guadiamar, Andalusia, southern Spain) Ferreras-Romero, Manuel; Cano-Villegas, Francisco J. & Rubio-Soler, M. Isabel
Departamento de Ciencias Ambientales (Zoologa), Universidad Pablo de Olavide, Ctra de Utrera km 1, 41013 Sevilla, Spain. <mferrom@upo.es>
In April of 1998, a dam that contained 6 hm3 of toxic sludge loaded of heavy metals, coming from mining activities, was broken. Pyritic sludge and acid water from the rupture in the dam of the Aznalcllar mining flooded 4,634 ha of the river Guadiamar basin. Toxic mud run 62 km, along river Agrio, reaching the river Guadiamar and arriving at the north of Gualdaquivir marshes. Important economic investments had been done by Spanish and Andalusian authorities in order to clean the sludge and minimize the serious ecological consequences caused in fauna and flora. Numerous scientific investigations are being done. It is intended to turn Guadiamar basin into ecological corridor, that connects protected areas of Guadalquivir marshes (Doana National Park) and Sierra Morena mountains (Sierra Norte Natural Park). Dragonfly communities of four reaches of the river Guadiamar are being studied: one is not affected by the mining accident (Gerena), two are affected by the accident (Sanlcar and Aznalczar), and one is affected as much by the mining accident as an industrial organic pollution (Quemas). Five criteria are being used: 1) number of species annually observed as adult (adult richness), 2) level abundance (according to Dufour 1978) of those species with upper number of records each year, 3) zoogeographical origin (St Quentin 1960, Ocharan-Larrondo 1988, Jacquemin & Boudot 1999) of most abundant anisopterans, 4) presence of focal species, they have been defined by us to this catchment (Platycnemis acutipennis, P. latipes, Cercion lindenii and Gomphus pulchellus), 5) taxonomic extent of use as breeding area of those reaches, number of species collected as larva or exuvia. Zygopteran richness and analysis with presence/absence data of focal species are useful criteria for the measure of the interannual change.
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28. Relationships between the concentrations of heavy metals in dragonfly larvae and Louro river sediments Rodrguez-Liares, G.1; Garrido, J.2; Bendicho, C.1; Lavilla, I.1
1 Departamento de Qumica Analtica y Alimentaria, Universidad de Vigo, Facultad de Qumica, As LagoasMarcosende s/n, 3200 Vigo, Spain. <isela@uvigo.es> 2 Departamento de Ecologa y Biologa Animal, Universidad de Vigo, Facultad de Biologa As Lagoas-Marcosende s/n, 3200 Vigo, Spain
Sediments provide a habitat for many aquatic organisms and these are a major deposit for most anthropogenic chemicals and waste materials, including heavy metals, that are introduced into surface waters. These heavy metals may be toxic for the biota, and their toxicity depends on their capacity of being incorporated to the food chain, becoming accumulated on the tissues. The bioavailable metal is the fraction that can interact with a biologic organism and be taken up to its structure. Bioavailabity of metals associated to sediments will depend on their solubility and their mobility. Nowadays, it is generally accepted that the environmental distribution, mobility and bioavailability of heavy metals not only depend on their total amount, but also are closely dependent on the association to the solid phase. The utilization of sequential extractions give a detailed information about the biological availability and the mobility of trace metals from the sediment. The sequential extraction is followed by using a sequence of reagents that release consecutively the different sediment fractions and metals associated with them (Filgueiras A. V., Lavilla I. and Bendicho C. 2004. Evaluation of distribution, mobility and binding behaviour of of metals in surficial sediments of Louro River (Galicia, Spain) using chemometric analysis: a case study. Science of the Total Environment, 330, 115 129). This work aims to study the relationships between the total amount of heavy metals in larvae of three species of dragonflies and the different fractions of sediments of the Louro river (tributary of the Mio river, Spain). The dragonfly larvae have a close contact with sediments; the species selected for the study were Cordulegaster boltonii (Donovan, 1807), Boyeria irene (Fonscolombe, 1838) and Onychogomphus uncatus (Charpentier, 1840). Total amount of As, Pb, Cd, Cr, Cu, Fe, Mn and Ni of dragonfly larvae were determined by inductively coupled argon plasma optical emission spectrometry (ICPAES). The sediments were analyzed by sequential extraction scheme BCR, that removes the following fractions: i) exchangeable ii) associated to carbonates iii) associated to Fe-Mn oxides iv) associated to organic matter and sulphides v) residual. The amounts of Pb, Cd, Cr, Cu and Ni in these fractions were determined by ICPAES. Major components of the sediments were determined by X-Ray Fluorescence.
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29. Evaluation of three species of dragonfly larvae as biomonitors of heavy metals Rodrguez-Liares, G.1; Garrido, J.2; Bendicho, C.1 & Lavilla, I.1
1 Departamento de Qumica Analtica y Alimentaria, Universidad de Vigo, Facultad de Qumica, As LagoasMarcosende s/n, 3200 Vigo, Spain. <isela@uvigo.es> 2 Departamento de Ecologa y Biologa Animal, Universidad de Vigo, Facultad de Biologa; As LagoasMarcosende s/n, 3200 Vigo, Spain
In general, the biomonitors are organisms which can be used for the quantitative determination of anthropogenically induced environmental factors. In aquatic ecosystems, benthic insects are the most frequently used biomonitors for heavy metals. There are, however, important unresolved problems in using aquatic insects as biomonitors: they require tedious sampling, sorting and identification to species. In particular, the small size of most of the insects is an important problem, because the determination of trace elements often requires sufficient biomass. On the other hand, there is a variability of the populations of the insects with the time and the locations. These technical problems are often overcome by studying the individuals grouped in genus, family or order, but that implies hiding a great amount of information, since different species may have different bioaccumulation, based on different uptake and growth rates. This work aims to evaluate different species of dragonfly larvae as accumulative biomonitors of heavy metals in Louro river. This river is one of the last tributaries of the Mio river, draining the largest part of the Valley of Louria, an area of 153 m2 with widespread human activities, mainly urban, industrial and agricultural ones. The species selected for the study were Cordulegaster boltonii (Donovan, 1807), Boyeria irene (Fonscolombe, 1838) and Onychogomphus uncatus (Charpentier, 1840), taking into account their close contact with sediments, their size, their wide distribution along the river and their ecological importance. Eleven sampling sites were selected along the course of the Louro river (points interval 2,5 km) affected by different anthropogenic sources of pollution. Total amount of As, Cd, Cr, Cu, Fe, Mn, Ni, Pb and Zn were determined by inductively coupled argon plasma optical emission spectrometry (ICPAES) after desiccation and acid digestion of samples. Pearson coefficients on metal accumulation were calculated to study differences on accumulation between the three different species, and principal component analysis (PCA) was applied to study accumulative associations between metals.
30. Dragonfly folklore in Galicia, Northwest Iberian Peninsula Fernndez-Martnez, Miguel A.

Tercio de Afora n9 Vigo, Spain. <miguelvacaloura@hotmail.com>
The objective of this project is to present a number of previously unknown names applied to odonata in Galicia, together with the associated belief system that, although apparently originating at different times in history, is nowadays fully coherent with the christian tradition. The data was collected between 1998 and 2004, each entry being accompanied with details of date and place of the recording and the name, age and birth place of the informant. The majority of informants (69%) were over 80 years old. 14 different names for adults and 3 for larval forms are recorded. They are grouped into six categories: those
78
alluding to the devil (Tizn do demo or 'Devil's ember', Cabalo do demo or 'Devil's horse', Cabalio do demo or 'Little devil's horse', Cabalo de San Vicente or 'St. Vincent's horse'); those alluding to the aggression relating to the eyes (Sacaollos and Quitaollos or remover of eyes); those alluding to similarity with other animals (Lavanco de catro alas or fourwinged duck, Grilo da auga or water cricket); those relating to shape and colour (gaiteiro or piper, candileiro and candil relating to oil-lamps); those relating to their usefulness to people (besbello and isca or fish-bait); and finally more recent names (libelia and liblula). People interviewed told of the danger of direct contact with odonates because of their link with the devil, as well as popular sayings about their lethal bite. Adults of the genus Calopteryx were seen used as bait when fishing for salmonids, as were the larvae of anisopteros caught emerging from mud taken from the bottom of pools. In Galicia no names were discovered that linked dragonflies with snakes as is common in other european cultures, but their bites are nonetheless considered lethal by some and snakes are thought by others to have originated from hair fallen into the water. Many Galician people still consider dragonflies to be dangerous animals into which the devil can transform himself, perhaps because of the large compound eyes at the front of the head and the pseudopupil, with the power of its gaze being a symbol of its wisdom. As such this is similar to the Cantabrian Caballucos del diablo or devils horses, seven large dragonflies allegedly ridden by seven demons, and characterised by their eyes that glow like embers. Names relating to damaging eyes refer to the danger of looking an odonate in the eye, as it is believed that one of the surest ways of being contaminated with the infamous mal de ollo or evil eye is the gaze of one of the devils servants or animals.
31. A new genus of Coenagrionidae (Odonata, Zygoptera) from the Pantepui region of Venezuela, with descriptions of four new species. de Marmels, Jrg
Universidad Central de Venezuela, Instituto de Zoologa Agrcola, Facultad de Agronoma, Apartado 4579, Maracay 2101-A, Venezuela. <demarmjc@hotmail.com>
The new genus includes seven species, all endemic in Pantepui (Guayana Highlands). Four are new to science, viz.: species nov. Nr. 1 from the Serrana de Maigualida, species nov. Nr. 2 from Cerro Yutaj and Cerro Yav, spec. nov. Nr. 3 and Nr. 4 both from Cerro Guanay. Aeolagrion chimantai De Marmels, 1988, Aeolagrion fulvum Needham, 1933, and Aeolagrion neblinae De Marmels, 1989 are transferred from Aeolagrion Williamson, 1917 to the new genus, which falls within Kennedys (1920, Ohio J. Sci. 21) Nehalennia-Telebasis series and shares characters with Telebasis Selys, 1865, and Aeolagrion flammeum (Selys, 1876). The most striking synapomorphy for all seven species is a fleshy or weakly chitinized basal ventral process on the male cercus.
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32. WAPODOKEY Mihokovic, Nino

Bribirska 39, HR-10000 Zagreb, Hungary. <nino.mihokovic@inet.hr>
The first key to species level of Croatian dragonflies was made using WAP (Wireless Application Protocol) language. The key is dichotomic, illustrated by the author and available on-line to users of mobile phones which support Internet connection. First check-list of Croatian dragonflies is given along with a brief description of species characteristics. It is available in Croatian, English, German and Spanish. To the authors knowledge this is the first and only scientific WAP key.
33. A comparative study of haemocytes in male and female of the dragonfly Crocothemis servilia Drury
Ashok, Jagtap1; Shinde, Kiran2 & Sathe, T.V.2

1 Vivekanand 2
College, Kolhapur, India. Dept. of Zoology, Shivaji University, Kolhapur - 416 004, India.
Haemocytes of both sexes of the dragonfly Crocothemis servilia Drury have been studied in this paper. Prohaemocyctes were in large mumber in both sexes. They were small, oval with smooth surface. The plasmocytes were marked by cytoplasmic processes. They were not polymorphic as compared to the proleucocytes. Spindle shaped haemocyctes were less in number compared to other cells. In both sexes proleucocytes, plasmocytes, sperule cells and oenocytes were observed. However spindle shaped cells were found less in number in males than in females. Oenocytes forms were segmentally arranged and occurred as clumps in the pleural epidermis near spiracles.
Abstracts Book 4th WDA International Symposium of Odonatology, Pontevedra 26-30 July 2005 Informal presentations
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Informal presentations Reflections on the 2003 International Odonatology Symposium, Beechworth, Australia Hawking, John H.
Murray Darling Freshwater Research Centre, Cooperative Research Centre for Freshwater Ecology, P.O. Box 991, Wodonga, 3690 Australia. <john.hawking@csiro.au>
The 3rd WDA International Symposiun of Odonatology was conducted at the historic township of Beechworth from January 8-13 2003. Beechworth is an old gold mining town set on the slopes of the western side of the dividing range in northeast Victoria. The setting for the symposium was the histroric old May Day Hills mental hospital, now part of La trobe University. Seventy five participants from 14 countries attended the symposium. Sixty papers were presented on a range of topic from ecology to taxonomy. The symposium was dedicated to three famous Australian odonatologists; R. J. Tillyard, A. OFarrell and J.A.L. Watson. The mid-symposium tour was a field trip from along the Kiewa River, from the mountains at Falls Creek to the plains at Kergunyah. The postsymposium tour was a four day tour, attended by 42 participants who visited native Koori people sites, historic towns, collected dragonflies and visited the famed bush ranger Ned Kellys region. The talk gives a pictorial account of the symposium.
Abstracts Book 4th WDA International Symposium of Odonatology, Pontevedra 26-30 July 2005 Author index
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Author index Alonso, A.................................................................................................................. 63, 67 Anselin, Anny................................................................................................................. 46 Ashok, Jagtap ................................................................................................................. 79 Azpilicueta, Mnica ....................................................................................................... 71 Baugne J-Y. .................................................................................................................. 44 Bedjani, Matja............................................................................................................. 37 Beirinckx, Kirsten........................................................................................................... 23 Bendicho, C. ............................................................................................................. 76, 77 Bhoje, P.M................................................................................................................ 61, 70 Bhosale, Y.A. ................................................................................................................. 56 Bogliani, Giuseppe ......................................................................................................... 33 Bouwman, Jaap............................................................................................................... 44 Buzatto, Bruno................................................................................................................ 62 Cano-Villegas, Francisco J. ...................................................................................... 71, 75 Carle, Frank L................................................................................................................. 52 Carvalho, Alcimar do Lago ............................................................................................ 55 Clausnitzer, Viola ........................................................................................................... 20 Contreras-Garduo, Jorge......................................................................................... 43, 62 Conze, K.-J. .................................................................................................................... 45 Corbet, Philip S. ............................................................................................................. 15 Cordero Rivera, Adolfo .............................................................. 26, 28, 34, 64, 65, 66, 71 Crdoba-Aguilar, Alex ....................................................................................... 29, 43, 62 De Knijf, Geert ............................................................................................................... 46 de Marmels, Jrg ............................................................................................................ 78 de Schaetzen R................................................................................................................ 44 de Vries, Henk ................................................................................................................ 74 Dvai, Gyrgy ................................................................................................................ 60 Dijkstra, Klaas-Douwe "KD" B.......................................................................... 20, 51, 53 Docampo Barrueco, Francisco ....................................................................................... 71 Dufrne, M. .................................................................................................................... 44 Fernndez-Martnez, Miguel A. ..................................................................................... 77 Ferreira, Snia .......................................................................................................... 68, 70 Ferreras-Romero, Manuel......................................................................................... 71, 75 Fichefet, V. ..................................................................................................................... 44 Fincke, Ola M. .......................................................................................................... 19, 23 Forbes, Mark................................................................................................................... 23 Fursov, Viktor................................................................................................................. 54 Garrido, J. ..................................................................................................... 63, 67, 76, 77 Giere, Sandra ............................................................................................................ 48, 65 Goffart, Ph. ..................................................................................................................... 44 Gonzlez de Castro, Ins ................................................................................................ 65 Gonzlez, A. ............................................................................................................. 63, 67 Graa, Manuel A. S. ....................................................................................................... 57 Groenendijk, Dick .......................................................................................................... 44 Grosso-Silva, Jos Manuel ....................................................................................... 68, 70 Gunasekaran ................................................................................................................... 57 Gnther, Andr ............................................................................................................... 31 Hadrys, Heike ............................................................................................... 20, 48, 63, 65
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Hawking, John H. ..................................................................................................... 38, 80 Hilfert-Rppell, Dagmar................................................................................................. 24 Holua, Otakar ................................................................................................................ 41 Kadoya, Taku ................................................................................................................. 33 Kalkman, Vincent........................................................................................................... 47 Karube, Haruki ......................................................................................................... 32, 56 Katatani, Naoharu........................................................................................................... 56 Kavane, P........................................................................................................................ 61 Ketelaar, Robert.............................................................................................................. 74 Khrokalo, Lyudmyla................................................................................................. 42, 72 Kitagawa, Kazuma ......................................................................................................... 56 Kjer, Karl M. .................................................................................................................. 52 Kurauchi, Yohei ............................................................................................................. 73 Lszl, Zoltn ................................................................................................................. 60 Lavilla, I. .................................................................................................................. 76, 77 Lebrun, Ph. ..................................................................................................................... 44 Leipelt, Klaus Guido ...................................................................................................... 52 Lorenzo Carballa, Olalla................................................................................................. 34 Marais, Eugene ............................................................................................................... 40 Margaj, G.S. ................................................................................................................... 56 Marinov, Milen............................................................................................................... 37 Martens, Andreas............................................................................................................ 36 Matthews, John H. .......................................................................................................... 50 Matushkina, Natalia.................................................................................................. 25, 72 May, Michael L. ....................................................................................................... 52, 66 McMillan, Victoria E...................................................................................................... 59 Mihokovic, Nino....................................................................................................... 62, 79 Mohan, Daniel ................................................................................................................ 57 Mundale, Mandar ........................................................................................................... 56 Nagy, H. Beta ......................................................................................................... 30, 60 Najera, Karla................................................................................................................... 62 Naskrecki, Piotr .............................................................................................................. 31 Ocharan, Francisco J. ..................................................................................................... 72 Olberg, Robert ................................................................................................................ 27 Olias, Marko ................................................................................................................... 37 Oppel, Steffen................................................................................................................. 69 Orr, Albert G. ................................................................................................................. 15 Ott, Jrgen ...................................................................................................................... 49 Pandharbale, A. R........................................................................................................... 61 Patil, R.G. ....................................................................................................................... 70 Paulson, Dennis .............................................................................................................. 18 Prez-Bilbao, A. ....................................................................................................... 63, 67 Plate, Calijn .................................................................................................................... 44 Prokopov, G.................................................................................................................... 42 Rey Muiz, Xos Carlos................................................................................................. 71 Rey Ra, Carlos............................................................................................................ 71 Riservato, Elisa............................................................................................................... 33 Rodrguez-Guntn, I.................................................................................................. 63, 67 Rodrguez-Liares, G. .............................................................................................. 76, 77 Rubio-Soler, M. Isabel ................................................................................................... 75
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Rppell, Georg ............................................................................................................... 26 Sahln, Gran ........................................................................................................... 17, 39 Sakoda, Tetsuo ............................................................................................................... 50 alamun, Ali ................................................................................................................... 37 Samways, Michael J. ...................................................................................................... 16 Snchez-Guilln, Rosa Ana................................................................................ 26, 64, 66 Sathe, T.V. .................................................................................................... 56, 61, 70, 79 Schenk, Kamilla ....................................................................................................... 27, 61 Schtte, Kai .................................................................................................................... 42 Serrano-Meneses, M. A. ................................................................................................. 29 Sherratt, Thomas............................................................................................................. 23 Shinde, Kiran ...................................................................................................... 61, 70, 79 Slavikovski, Ana............................................................................................................. 62 Soares-Vieira, Patrcia .............................................................................................. 68, 70 Sousa, Pedro ................................................................................................................... 68 Spector, Sacha ................................................................................................................ 31 Suda, Shin-ichi ............................................................................................................... 33 Suhling, Frank .......................................................................................................... 36, 39 Susa, Koichi.................................................................................................................... 35 Szab, Zoltan D. ............................................................................................................. 30 Szllassy, Nomi ...................................................................................................... 30, 60 Szkely, Annamria........................................................................................................ 60 Szkely, T. ...................................................................................................................... 29 Taylor, Philip D. ............................................................................................................. 18 Termaat, Tim .................................................................................................................. 74 Thite, H.S........................................................................................................................ 70 Thomas, Manu ................................................................................................................ 57 Thompson, David J......................................................................................................... 21 Timm, Janne ................................................................................................................... 63 Torralba Burrial, Antonio ............................................................................................... 72 Tsubaki, Yoshitaka ................................................................................................... 33, 48 Ubukata, Hidenori .................................................................................................... 50, 73 Van Gossum, Hans ............................................................................................. 23, 64, 65 Wargel, Antonia.............................................................................................................. 65 Washitani, Izumi............................................................................................................. 33 Watanabe, Mamoru .................................................................................................. 22, 35 Watanabe, Yoko ....................................................................................................... 37, 74 Watts, Phillip C. ............................................................................................................. 21 Weihrauch, Florian ......................................................................................................... 37 Worthington, Andrea ...................................................................................................... 27 Yadav, R. P..................................................................................................................... 61
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LIST OF PARTICIPANTS
Surname Ackerman Alonso Prez Arnold Ashok Averill Ayres Fernndez Bernard Bouwman Bower Cano Villegas Carvalho Claustnizer Conze Corbet Cordero Rivera Crdoba Aguilar De Knijf DeMarmels Dijkstra Domnguez Borines Domnguez Borines Dow Fernndez Martnez Ferreira Name Jennifer Ana Isabel Rob Jagatap Mike & Linda Csar Rafal Jaap Jennifer Francisco Jess Alcimar Viola KlausJuergen Philip S. Adolfo Alejandro Geert Jrg KlaasDouwe Jessica Vernica Rory Miguel A. Organisation National Geographic Society University of Vigo City Country Charlottesville USA Vigo Spain email ackerman@cstone.ne aiap@uvigo.es RArnold@mail.colgate.edu
Colgate University Hamilton USA Vivekanand Kolhapur India College Kidderminster UK University of Vigo Adam Mickiewicz University Dutch Butterfly Conservation Universidad Pablo de Olavide Universidade Federal do Rio de Janeiro Philipps University of Marburg Pontevedra Pozna Wageningen Pen Selwood Sevilla Rio de Janeiro Halle Essen University of Edinburgh University of Vigo Instituto de Ecologa, UNAM Institute of Nature Conservation Universidad Central de Venezuela Natural History Museum Leiden St Buryan Pontevedra Mxico D.F. Brussels Maracay Leiden Vigo Vigo Spain Poland
miketaverill@aol.com Lindamaverill@aol.com, cesar_ayres@yahoo.com rbernard@amu.edu.pl
The jaap.bouwman@Vlinderstichting.nl Netherlands UK bowerje@btopenworld.com Spain fjcanovi2@hotmail.com Brazil Germany Germany UK Spain Mxico Belgium Venezuela alagoc@acd.ufrj.br violacl@gmx.de Klaus-juergen.conze@t-online.de mail@pcorbet.vispa.com adolfo.cordero@uvigo.es acordoba@ecologia.unam.mx geert.deknijf@inbo.be demarmjc@hotmail.com
The dijkstra@nnm.nl Netherlands Spain galatea_8@hotmail.com Spain tucuxi4@hotmail.com rory.dow@virgin.net miguelvacaloura@hotmail.com
Snia
Coulsdon UK Ecoplanin, Vigo Spain Xestin e Informacin Ambiental CIBIO-Univ. Porto Vila do Conde Portugal
hiporame@gmail.com
85
Surname Ferreras Romero Fincke Fitch Fursov Garca Moreno Marchn Garrido Giere
Name Manuel Ola David Allan Viktor Josep Josefina Sandra
Organisation Universidad Pablo de Olavide University of Oklahoma National Academy of Sciences
City Sevilla Norman Lexington Kiev Barcelona
Country Spain USA USA Ukraine Spain Spain Germany
email mferrom@upo.es fincke@ou.edu DAF@fitchlawoffices.com vfursov@ukrpack.net jgarciamm@terra.es jgarrido@uvigo.es sandra.giere@ecolevol.de
University of Vigo
Tierrztliche Hochschule Hannover
Vigo Hannover
Goffart
Philippe
Gonzlez de Ins Castro Graa Manuel Grosso-Silva Jos Manuel Gnther Andr Hadrys Hawking HilfertRppell Hoffmann Holua Honkavaara Heike John Dagmar Joachim Otakar Johanna
Centre de Gembloux Recherche de la Nature, des Forts et du Bois University of Vigo Pontevedra Universidade de Coimbra CIBIO-Univ. Porto TU Bergakademie Freiberg University of Hannover Murray Darling Freshwater Res. Center Coimbra
Belgium
p.goffart@mrw.wallonie.be
Spain Portugal
inesglezdecastro@yahoo.es mgraca@ci.uc.pt jmgrossosilva@yahoo.com andre.guenther@ioez.tu-freiberg.de Heike.Hadrys@ecolevol.de john.hawking@csiro.au rueppell-film@t-online.de joa.hoff@t-online.de holusao@email.cz johonkav@bytl.jyu.fi marcianoh@tiscalinet.it kadoya@e-mail.jp
Vila do Conde Portugal Grossschirma Hannover Albury Cremlingen Germany Germany Australia Germany Germany Czech Republic Finland Italy Japan
alauda Mendel University of Agriculture and Forestry University of Jyvskyl The University of Tokyo RMNH Leiden
Hamburg FrydekMstek Jyvskyl Vercelli Tokyo Leiden
Huancahuari Marciano Kadoya Taku Kalkman Karube Khrokalo Kurauchi Vincent Haruki Lyudmyla Yohei
Kanagawa Odawara Prefectural Museum Kyiv National Kiev Taras Shevchenko University University of Tsukuba Tsukuba
The kalkman@nnm.nl Netherlands Japan qtmfc457@ybb.ne.jp Ukraine Japan lkhrokalo@mail.ru s0520231@ipe.tsukuba.ac.jp
86
Surname Lszlo Leipelt Lorenzo Carballa Marais Martens Matthews Matushkina May McMillan Mihokovic Mill Mundale Nagy
Name Zoltn Klaus Guido Olalla Eugene Andreas John Natalia Michael L. Victoria Nino Peter Mandar H. Bata
Organisation University of Debrecen Technische Universitt Braunschweig University of Vigo National Museum of Namibia University of Education University of Texas, Austin Kiev National University Rutgers University Colgate University
City Debrecen
Country Hungary
email laszlozoltan@gmail.com k.leipelt@tu-bs.de olalla.lorenzo@uvigo.es insects@natmus.cul.na martens@ph-karlsruhe.de johoma@mail.utexas.edu matushkina@list.ru mimay@rci.rutgers.edu vmcmillan@mail.colgate.edu nino.mihokovic@inet.hr gpmill@supanet.com mandar.mundale@gmail.com nagy.beata@gmail.com cdwadei_2004@yahoo.com
Braunschweig Germany Pontevedra Windhoek Karlsruhe Austin Kiev New Brunswick Hamilton Zagreb Leeds Kolhapur Debrecen Kathamdnu Spain Namibia Germany USA Ukraine USA USA Croatia UK India Hungary Nepal
Shivaji University University of Debrecen Nepali Yam Centre for Drinking Bahadur Water, Agriculture Development and Environment Improvement Norling Ulf Malmo University Orr Albert G. Griffith University Ott Jrgen L.U.P.O. GmbH Pandharbale Ashok Ram Pandharpur College Pardo Isabel University of Vigo Gamundi Patil Ramarao L.B.S. College Gunda Satara Paulson Dennis University of Puget Sound Peitzner Gabi Pritchard Gordon University of Calgary Reels Graham Asia Ecological Consultants Ltd Rey Muiz Xos Lois Sociedade Galega de Historia Natural Riservato Elisa University of Pavia Rodrguez Iria University of Vigo Guntn Rowe Richard James Cook University
Lund Brisbane Trippstadt Pandharpur Vigo Satara Tacoma Boernsen Calgary Yuen Long Vilagarca Pavia Vigo Townsville
Sweden Australia Germany India Spain India USA Germany Canada Hong Kong Spain Italy Spain Australia
ulf.norling@ts.mah.se agorr@bigpond.com L.U.P.O.GmbH@t-online.de
ipardo@uvigo.es
dpaulson@ups.edu gabi.peitzner@gmx.de gpritcha@ucalgary.ca gtreels@asiaecol.com.hk xoselois.rey@terra.es elisa.riservato@unipv.it iriaguntin@uvigo.es richard.rowe@jcu.edu.au
87
Surname Rubio Soler Rppell Sahln Samways Samways Snchez Guilln Sathe Schenk
Name M Isabel Georg Gran Melinda Michael Rosa Ana
Organisation Universidad Pablo de Olavide Halmstad University University of Stellenbosch University of Stellenbosch University of Vigo
City Sevilla Cremlingen Halmstad Matieland Matieland Pontevedra Kolhapur
Country Spain Germany Sweden South Africa South Africa Spain India Germany
email mirubsol@upo.es rueppell-film@t-online.de goran.sahlen@set.hh.se
samways@sun.ac.za rguillen@uvigo.es pmb_bhoje@yahoo.co.in k.schenk@tu-bs.de
Tukaram V. Shivaji University Kamilla Institut fr
Geokologie, TU Braunschweig
Darmstadt Hamburg Bath Kolhapur Purley Zagreb Rio Maior Germany Germany UK India UK Croatia Portugal USA Germany Jsilsby1@btinternet.com ana_slavikovski@net.hr pmsvieira@yahoo.com spector@amnh.org f.suhling@tu-bs.de w.schneider@hlmd.de Brancsikia@web.de bspmasm@bath.ac.uk
Schneider Schtte Serrano Meneses Shinde Silsby Slavikovski SoaresVieira Spector Suhling
Wolfgang Kai Martn Alejandro Kiran Pandurang Jill Ana Patrcia Sacha Frank
Hessisches Landesmuseum University of Hamburg University of Bath Shivaji University
Susa Szabo Szallassy Takahashi Taki Taylor Termaat Thomas Thompson Timm Torralba Burrial
Koichi Zoltan Daniel Noemi Yuma Wakana Philip Tim Manu David J. Janne Antonio
American Museum of Natural History Institut fr Geokologie, Technische Universitt Braunschwei University of Tsukuba Babes-Bolyai University Babes-Bolyai University University of Tsukuba University of Tsukuba Acadia University Dutch Butterfly Conservation Madras Christian College University of Liverpool University of Oviedo
New York
Tsukuba Cluj Cluj Tsukuba Tsukuba Wolfville Wageningen Chennai Liverpool Bremen Oviedo
Japan Romania Romania Japan Japan
susa974@hotmail.com szabodz@hasdeu.ubbcluj.ro szallassy@gmail.com yuyuyuyu@mail2.accsnet.ne.jp s0310780@ipe.tsukuba.ac.jp
Canada philip.taylor@acadiau.ca The Tim.Termaat@Vlinderstichting.nl Netherlands India mtmathai@hotmail.com UK Germany Spain D.J.Thompson@liverpool.ac.uk janneket@gmx.de antoniotb@hotmail.com
88
Surname Tsubaki
Name Yoshitaka
Ubukata
Hidenori
Van Gossum Hans Velo Antn Wain Wain Wargel Guillermo Chris William Antonia
Organisation National Institute for Environmental Studies Hokkaido University of Education University of Antwerp University of Vigo British Dragonfly Society British Dragonfly Society
City Tsukuba
Country Japan
email tsubaki@nies.go.jp
Kushiro Antwerp Pontevedra Bordon Bordon Hannover
Japan Belgium Spain UK UK Germany
ubukata@kus.hokkyodai.ac.jp Hans.VanGossum@ua.ac.be guillermov@uvigo.es TheWains@ukonline.co.uk TheWains@ukonline.co.uk A.Wargel@gmx.de
Tierrztliche Hochschule Hannover

University of Tsukuba
Watanabe Watanabe Weihrauch Worthington Yadav
Mamoru
Tsukuba Nishinomiya Wolnzach Niskyauna Ichalkaranii
Japan Japan Germany USA India
watanabe@kankyo.envr.tsukuba.ac.jp mer-yoko.watanabe@nifty.com Florian.Weihrauch@t-online.de WORTHINGTON@siena.edu
Yoko Florian Andrea Siena College Ramchandra D.K. A.S.C. P. College Ichalkaranii
Funding agencies:
Universidade de Vigo
Ministerio de Educacin y Ciencia

Program Abstracts 4th WDA Symposium PDF

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Abstracts Book 4th WDA International Symposium of Odonatology, Pontevedra 26-30 July 2005

Program and Abstracts

Edited by Adolfo Cordero Rivera, University of Vigo

Corbet, P.S. Orr, A.G.

Sexual selection Monitoring and Spector, S. climate change

A Po bra d o car ami al

Vila no va de A rou saV ilan ova d e A

COM P LEX O HM IDO D E CO RRUB E

16:30 17:30 17:45 18:00 18:15 18:30 20:45

17:30 17:45 18:00 18:15 18:30 19:45 ??

15:45 16:00 16:15 16:45

16:00 16:15 16:45 17:15

Sunday 31 July to Thursday 4 August, Postsymposium tour

List of poster presentations

Corbet, Philip S....................................................................................................................................................15

Movement behaviors of odonates in heterogeneous forest landscapes..................................

Hadrys, Heike ......................................................................................................................................................20

Flashes in flight communication between odonate males .....................................................

Serrano-Meneses, M. A., Szkely, T. and Crdoba-Aguilar, Alex ................................................................29

Szllassy, Nomi, Szab, Zoltan D., Nagy, H. Beta......................................................................................30

Why are endemic Odonates endangered in oceanic islands Ogasawara?................................

Dragonflies of riverine habitats : assessment as indicators of biodiversity and environmental integrity...................................................................................................................

Riservato, Elisa & Bogliani, Giuseppe..............................................................................................................33

Ecology of Odonata inhabiting permanent Namibian desert springs.....................................

Hawking, John H.................................................................................................................................................38

Suhling, Frank & Sahln, Gran .......................................................................................................................39

Marais, Eugene ....................................................................................................................................................40

Contreras-Garduo, Jorge & Crdoba-Aguilar, Alex.....................................................................................43

The Dutch Dragonfly Monitoring Scheme: results and trends. ...............................................

Tsubaki, Yoshitaka ..............................................................................................................................................48

Giere, Sandra & Hadrys, Heike .........................................................................................................................48

Thomas, Manu, Gunasekaran and Mohan, Daniel .........................................................................................57

Graa, Manuel A. S. ............................................................................................................................................57

Mihokovic, Nino & Slavikovski, Ana ...............................................................................................................62

Timm, Janne & Hadrys, Heike ..........................................................................................................................63

Wargel, Antonia, Giere, Sandra & Hadrys, Heike...........................................................................................65

May, Michael L. ...................................................................................................................................................66

Snchez-Guilln, Rosa Ana & Cordero Rivera, Adolfo.................................................................................66

Present knowledge on the Odonata of Serra da Estrela Natural Park (Portugal)....

Biodiversity of dragonflies (Odonata) from Western ghats of Maharashtra, India .

Cano-Villegas, Francisco J. & Ferreras-Romero, Manuel..............................................................................71

Flight peak trends for dragonflies from the Netherlands ............................................

Morphological characteristics of odonate eggs and early instar larvae specific to

Watanabe, Yoko ............................................................................................................................... 74

Ferreras-Romero, Manuel; Cano-Villegas, Francisco J. & Rubio-Soler, M. Isabel.....................................75

Evaluation of three species of dragonfly larvae as biomonitors of heavy metals ....

Dragonfly folklore in Galicia, Northwest Iberian Peninsula.......................................

Author index.................................................................................................................................... 81 LIST OF PARTICIPANTS .......................................................................................................... 84

Hawking, John H.................................................................................................................................................80

Oral contributions Forests as habitats for dragonflies (Odonata) Corbet, Philip S.

Movement behaviors of odonates in heterogeneous forest landscapes Taylor, Philip D.

The importance of forests to neotropical dragonflies Paulson, Dennis

Documenting odonate behaviour by drawing from films Hilfert-Rppell, Dagmar

Ovipositor and egg-laying behaviour of Odonata: phylogenetic implications Matushkina, Natalia

Flashes in flight communication between odonate males Rppell, Georg

Sperm ejection as a cryptic female choice mechanism in odonates Crdoba-Aguilar, Alex

The global dragonfly assessment Spector, Sacha1 & Naskrecki, Piotr2

Artificial parthenogenesis in Aeshna nigroflava Martin Watanabe, Yoko

Dragonfly monitoring in northrhine Westfalia, Germany Conze, K.-J.

Towards an atlas of European odonates Kalkman, Vincent

Ecomorphology of legs in larval and adult Anisoptera Leipelt, Klaus Guido

30. Dragonfly folklore in Galicia, Northwest Iberian Peninsula Fernndez-Martnez, Miguel A.

32. WAPODOKEY Mihokovic, Nino