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1A ZaraTabi(ztabi@ucsd,edu) BICD 120:Dr.

Schmidt
AIM:ztabi06 Week2
Details
TheNitty-Gritty
Photosynthesis:
+ 60z + 12NADP*+18ADP+ 6 HzO
6COz+ 12HzO+12NADPH+ 18ATP) CoHrzOo

Production
TheLiqhtReactions: of and fromlightenergy!
inPhotosynthesis
Pigments
- lmportant
. Primary pigment;
photosynthetic ) located
inthereaction ofeachphotosystem
center
. Secondary/Accessory Help
Pigments; spectrum byplant
of lightthatcanbeabsorbed
inthe
) located (LHC)orAntennaComplex

. Beableto interpret
graphs andActionSpectra!
oftheAbsorption
- Understandwhatwasconcluded T.W.Engleman's
from experiment

unit=
-Photosynthetic
. Sizeof LHChelpsprevent bycontrolling oflightabsorbed
theamount
isthecauseof photooxidation
whichforms
cell-damaging

toprevent
abletogatherfreeradicals damage
. Growing
in ) sizeof LHCtoabsorbmorelightenergy
) BIARatio
chlorophyll
. Growing
in ) lightintake
sizeof LHCtoregulate
) chlorophyll
B/Aratio

ofeachprotein
- Knowthenameandlocation complex:
6trorna- th t
NADP}T

ligvtt ztl+
vi
?\r-,Pf'notto

2$+
9*o' PlooaY'wiru-
[r
Vs.Noncyclic
- Cyclic Photophosphorylation
. lnNon-Cyclic
photophosphorylation:
- UseofBOTH Photosystem ll andI
of 2HzOproduces e-,-
- Splitting Oz,afld NADPH
H*pumped intothylakoidlumenper2HzO) _ viaphotophosphorylation
ATPproduced
quanta
oflighttotaltomove4 e-) - per photosystem
each
. InCyclicphotophosphorylation,
no isavailabletobereduced, backto
soe-areshuttled
topumpmoreH+) MoreATPProduced
- Uses onlyPhotosystem ) NoHzO issplit

TheCatvin Photosvnthesis-
Cvcle/DarkReactions/C3 fromCOzFixation
foodsource
Producing
by
- Discovered
- COzinitiallyfixedintoa 3-Cmolecule(

@co' 0sbisco
- o GCin+efinediate
@ ribulo$obiph ospvratu \
3rc0<-f- feuor;
q @) O-piorpl,ogrScrrcre-
DA-
r' 'rP
a#toororrADPll aN+
O GOP ).^

ffi ';lHil],5;
,?, - a,4
1#:,s,a,,che
- Need "
ATPand
withRubisco
- TheProblem
Ozlandt
NADPHPerCO2Fixed
forBOTH
) hasanaffinity
COz]= Rubisco to
binds
and-
'[ -
. L-Ozland t COzl= Rubisco to
binds

method
C4Photosynthesis-Another ofcarbonfixation,
discovered HatchandSlack(1965)
byscientists
- WhyCalledC4?) COzFixedendsupina 4-Cmolecule( insteadof3-C!
- C4andC3photosynthesis
are
C.,$t
--*llrCo3
llzD (Ltc)
PTP NA,DP+NAppt{
- ctt
Cetts
ilesoghgtl
n4alate +!- O,tn

v\alqYg
P!r uv aw4
,/

-Whydoit whenit usesupmoreenergy (intheformof highenergyphosphates,


whenATp) AMp)?
. Makes a PROCESS:
1) hasa higheraffinity
forCOzincomparison
to
2)Create anenvironment of tofavorCO2fixation
overphotorespiration
3)Arrangement ofcells(C4or KranzAnatomy) helps thatmay
havebeenlostduring photorespiration

- Because
CO2loss
is stomata
moreoften) lessH2Oloss!
. ldealforplantsthatliveinareasof high and
whererateofphotorespiration
increases
ex.TropicalGrasses-Maize,
Sugar Cane,
Millet,
etc.
CAM(Cfas$ulacean
AcidMetabolisml
Photosvnthesis-
Analternate
version
ofC4photosynthesis
- C4andC3Reactions
are_ separated-Day
andNiqht

Cot
+ Hrco,
Fzo

ATP

- Becausehigh[] of (malicacid)storedinthevacuole
atnight,pHwillusually
be
- Whydiditevolve?
. Helpplantsthatlivein locations
withlittle
- Keepstomata duringthedaytimetopreventwaterloss
) Canstillperform
C3photosynthesis
stomataatnightwhentemperatureiscoolertotakeinco2 toperform
c4
photosynthesis
. ex.Cacti,
succulents,pinepple

PrimaryProductivitv,
GlobalWarminq-
Questions tothinkabout:
- Definition
forprimary
productivity?
- Whattypesofplants(C3,C4,CAM,etc.)arethemostefficient
atfixingCO2?Leastefficient?
- Whatcontinental playsanimportant
ecosystem roleinfixing
themosttarbonperyear?
- HowmanyPounds ofCO2arereleasedforevery gallonofgasused?
(Refer
tothehandout givenon10/08/09
lecture!)