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ABSTRACT. Live fence posts are widely used in Costa Rica and
other Cenhal American countries as a sustainable agricultural prac-
tice. The Costa Rican farmers' empirical knowledge was compiled
through a questionnaire, field measurements and literature review.
Ninety two species used for live fences were recorded and tabulated.
The management practices (preparation of stakes,planting, attaching
wire, pruning regimes) and the various uses are discussed. The bio-
mass production of a kilometer of one live fence was measured. It is
concluded that this is indeed a very promising sustainable practice
which deserves more research and dissemination.
INTRODUCTION
cuttings or stakes (about 2-2.50 m). Each species has its own particular
growth characteristics, cultivation requirements, management practices,
derived products, and benefits. These trees may provide wood for fuel and
charcoal or for construction (poles, posts, pillars, etc.), edible fruits and
flowers, flowers for honey, leaf forage for cattle and other domestic mi-
mals (e.g., goats, rabbits, and chickens), handicraft (seeds used for beads,
ornamental wood), medicinal products, gums and resins, dyes as well as
various other products. These plants also have great value as ornamentals
and as a refuge for wild animals, notably birds. Additionally, they continu-
ally yield new cuttings for more fence posts. However, their main purpose
is to serve as physical support to attach rows (usually 3, but their number
may vary from 2 to 5) of barbed wire so as to effectively protect houses,
crops, cattle and pastures or differenttypes of gardens. Furthermore,living
fences provide conspicuous and fm boundaries to separate properties or
areas within properties or pastures. When cuttings are planted very closely
and sometimes associated with smaller plants that may be spiny (e.g.,
Bromelia pinguin), or poisonous (e.g., Euphorbia cotinifolia and Hura
crepitans), they can also provide an effective barrier to cattle and people,
even without use of barbed wire.
Many other benefits and services are also provided. The leaves shed by
live fences serve as mulch and release nutrients to adjacent crops. The
periodic pollarding (cutting back the crown of the fence posts) results in
starving and death of roots, leading to small air channels in the soil which
favors water infiltration. Other benefits include nitrogen fixation by some
species, specially legumes, erosion control and better infiltration of water,
provision of shade, use as wind-breaks, niches for insecteating buds,
support for orchids and ornamental plants and many other additional uses.
A fair statement is that they have considerableaesthetic value. Whether as
part of the landscape or from the beauty of their flowers and branching
forms, live fence rows provide a pleasant break to the monotony of many
grass-dominated landscapes.
Many farmers, ranchers and rural laborers are experienced with the use
of live-fence posts; some have well defined criteria concerning planting
practices and the advantages and disadvantages of various species. But
their knowledge is empirical and with the exceptionof a few species, such
as Gliricidia sepium (Baggio, 1982; Beliard, 1984), this has not been
quantified. The purpose of the present investigation is to overview present
technologies and beliefs concerning live fence posts used in Costa Rica
and to support obse~atiofl~ with farmer interviews, some field measure-
ments, a literature review, and other experiences by the authors over the
last 10 years.
68 JOURNAL OF SUSTAINABLE AGRICULTURE
METHODS
1. For convenience,here the word lowland applies from sea level to about 600m
elevation, middle elevation from 600 to 2000 m and highland further above. The
limits of 600 and 1500 represent approximately the lower and higher ranges
adequate for commercial coffee cultivation.
Research. Reviews, Practices and Technology 71
The harvesting of stakes for more posts always takes place from the top
of the trunk (originally a cutting itself). The top gradually assumes a
swollen bulb or headlike appearance (callus). Owing to the propensity of
fence mee species to sprout vigorously with many new shoots after pol-
larding, the upper part of the stake may be much larger than the diameter
of the stake itself. This impression may be increased by the abundant
growth of epiphytes that find suitable niches between the branches or the
stubs of past pruned branches. Cutting back repeatedly promotes extensive
callus and woody tissue that progressively build up from wound healing.
A typical fence post of Erythrina costaricensis, for example has a trunk
top diameter between one and a half and two times that of the trunk.
Preparation ofposts: The majority of the important fence forming trees
in Costa Rica are propagated by planting large stakes (cuttings), usually
around 4-12 cm diameter at the base, and 1.5-2.5 m in length, depending
on the purpose of the fence and the preferences of the farmer. Cuttings are
derived from branches arising from live fences that are pollarded after one
or two years of growth. The number of shoots produced by each estab-
lished live fence tree varies according to age, vigor and period since the
last pollarding. A living post can normally produce from 2 to about 8
cuttings for planting every two years. Cuttings are easy to prepare and
require no special skill in planting, although handling with care between
cutting and planting is necessary. From early February through early
April, mostly when the weather is relatively dry, the trees are pruned to
obtain cuttings for planting as well as to remove excess or undesirable
growth from the parent tree. Many of the unused branches and leaves form
a mulch around the fence. However, cutting has also taken place at other
periods during the year.
The time-lag between the planting of a cutting and when it begins to
produce shoots for new cuttings of plantable size varies among species but
it is generally six months to two years. Sometimes it is even less as in the
case of the fast growing Erythrina spp. and Gliricidia sepium. Once this
production is initiated, sprouting vigor apparently does not decrease for a
relatively long time (over 20 years, at least, has been recorded for some
well known species with no apparent decline). When in healthy conditions
the trees show considerablelongevity. Older trees of Gliricidia sepium can
display deformationwhere the barbed wire has been attached. Some fence
trees have been estimated to be 90 years old. However, species with
relatively soft wood such as Bursera sirnaruba,and Spondias purpurea are
reported by local farmers to have a shorter life expectancy than species
with denser wood, such as Gliricidia sepium or Diphysa robinioides.
The annually repeated pollarding seems to enhance juvenility. The
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Research, Reviews. Practices and Technology 75
woody shoot regrowth of live fence trees generally appears to have more
leaves in proportion to the amount of woody tissue than shoots of naturally
growing trees of similar age. They are also said to show higher rooting
success than shoots from non pollarded trees. However replicated tests are
needed to c o n f i i these differences.
Erect, straight, vertical branches are preferred over crooked or lateral
branches. Both are found along fences, sometimes side by side. But it is
also common to F i d entire rows of straight vertical branches, a possible
indication that the same clonal material had been used. Accordine to some
farmers, the progeny of crooked cuttings manifests the curving teidencies,
making for less even fences. Lateral branches reputedly have lower root-
ing effectiveness than more vertical branches. Both these beliefs need to
be confiied.
From the last week of January, throughout February, March and into
mid-April, considerable activity along the fence lines takes place, as
shown by abundant evidence: pollarded pruned fence rows, leafy material
on the ground, stacks of smaller branches piled for fuelwood, cuttings
placed under some shade, with lateral branchlets and leaves neatly re-
moved to produce "clean" poles and of course newly set live fence posts.
The coincidence of this seasonal silviculturalpractice with the major dry
season h u g h o u t Costa Rica is considered quite important. The major
factors in the efficacy of this timing seems to be related to the effect of the
approaching rains and morpho-physiologic periodicity of the species. In
the dryer areas, many cuttings are leafless or have few leaves. Even in the
wetter areas, there seem to be less leaves at that time. The practices which
are followed are predicated upon the fact that planting should precede the
onset of the heavy rains so that root and bud initiation will already have
begun when the season starts.
The cuttings are prepared before the appearance of buds that precede
the vegetative growth flush associated with the rainy season. Flowering
cuttings are reputedly not good propagation material. Even in locations
with adequate soil moisture it is said that lower percentage rooting success
is obtained when cuttings are derived from flowering branches.
It is usually considered unadvisable to plant the cuttings during the
rainy season itself, but many exceptions exist. The reasons given for no
planting in the rainy season are the interruption of active vegetative
growth, lack of time for adequate healing of cuttings and subsequent
susceptibility to fungal infections due to excessive water and poor soil
drainage which limit the production of roots. The seasonal changes in
rooting effectiveness may involve carbohydrate mobility, hormonal fac-
tors and bud dormancy, all of which need to be investigated.Many farmers
76 JOURNAL OF SUSTAINABLE AGRICULTURE
as a backing, thus enabling the wire to move above the bark and not to
penetrate into it.
Other management practices: During the first year, leafy side branches
from the trunk are carefully pruned off. This practice serves to direct
growth to the top shoots and to train the tree by promoting corollary
inhibition of lateral shoot initiation from the trunk. At the same time, any
undesirable lateral, weak, or excessive top branches are removed.
The gaps between the living fence posts are gradually filled with new
cuttings to reach the desired density. Densities range from 1to 3 posts per
meter up to 6 posts per meter in very dense fences around coffee and
vegetable fields and gardens to prevent animal entry as well as to protect
against strong winds. Most common is a density of 2-3 trees per meter.
Another variation is the planting of widely spaced, well-rooted posts 4-5 m
apart, with living posts planted in between. The latter may or may not be
attached with barbed wire. In dry areas the spiny Bromeliapinguin (which
has edible fruits) is a common candidate to fill in the lower portion of the
fence.
Cuttings for reinforcement usually are of smaller diameter than those
used for the establishment of the original fence. The cuttings are some-
times tied to the barbed wire to train them during the period preceding
nailing of the barbed wire. W~ththe planting of new cuttings to fill in an
older fence, other welldeveloped fence trees are sometimes harvested for
firewood, or for construction posts and occasionally lumber and cabinet
wood This practice allows a gradual replacement and harvesting without
depleting the area of support for barbed wire.
Shchez and Russo (1985) reported preliminary results of a survey by
questionnaires carried out in Costa Rica by the Nitrogen Fixing Trees
Project (NFTP), CATIE, Tunialba on uses of Erythrina spp. including
management practices. Table 2 details the findings in relation to fence
posts. Later, Shchez and Payne (1987) communicated results of another
survey done by the NFTP on Gliricidia sepium. The authors found that
95% of the interviewedfarmers use the species as live fence posts.
Uses: For most species both the main stem and the cuttings produced
by branches provide a much needed supply of firewood,although the latter
is considered of lesser quality since insufficient time is allowed to produce
heartwood. The quality varies by species, those with the highest density a
T a b l e 2. Hanagement p r a c t i c e s o f E r y t h r i n a s p p . f o r f e n c e p o s t s
I n C o s t a R l c a (Sanchez a n d Russo, 1 9 8 5 ) .
E. b e r t e r o a n a 50 % Large c u t t i n g s 96 %
2. fUSCd 5 % Direct planting by seed 4 %
Other Erythrinas. 45 %
Diagonal 10 % Diagonal 82 %
Tvo f a c e s 10 % ConicaI 4 4
Three faces 10 % Flat 14 %
Conical 19 %
Flat 51 %
Up 7 days 18 % Dry p e r i o d 35 %
8 t o 15 days 41 % S t a r t of rainy season 22 %
Without r e s t 41 % During t h e rainy season 22 %
Throughout t h e y e a r 17 %
MOON ?SASE
Rising 0 - 1 m
Wanin? 1.1 - 2 . 0 m
2 . 1 - 3.0 rn
3.1 - 4.0 m
Variable
< 49 % 6 % NO
50 -
69 % 12 % Yes
1 0 - 79 '1 23 %
8 0 -. 8 9 % 6 %
90 -100 % 53 %
. .
the American tropics, yields a wood of similar qualities but its diameter
growth is slower, although it produces cuttings at a faster rate. It has
durable wood which is used for cabinets, for artisanal work and for poles
in contact with the ground. A workshop on management and improvement
of this species was recently held in the Tropical Agricultural Center for
Research and Training (CATIE), Turrialba,Costa Rica (Withington et al.,
1987).
few other fence trees are often allowed to grow into adult trees
because of the value of their wood. Among those Bombacopsis quinata,
Tabebuia rosea and Tectona grandis stand out in the lowlands. In the
highlands, Cupressus Iusitanica and Alnus acuminata, both propagated by
seed, are most common.
The leaves, flowers and fruits of several live fence trees are an impor-
tant source of fodder for domestic animals and food for humans. Little
research on this subject has been done, especially in the lowland humid
tropics. The leaves of Gliricidia sepium are commonly fed to cattle, but
are said to be semi-toxic to horses. The flowers, however, are edible when
cooked.
Spondias purpurea, a cosmopolitan tree, bears red or yellow plum-like
edible fruits appreciated by livestock as well as humans. The leaves are
also eaten when young. Some varieties, particularly with large fruits are
often seen in markets. The tree shows surprising variation in appearance
and growth form. There is great variation in the quantity and quality of
fruit, but the highest yields and the best quality come from rather dry
areas. The names of some particularly productive varieties have been
reported from Guatemala (Standley and Steyermark, 1946).
Eugenia jumbos bears a crisp fruit, the rose apple and is grown from
polyembryonic seeds which produce multiple fast growing shrub-like
stems. The tree is often planted for windbreaks to protect coffee planta-
tions and other crops at middle elevations. Erythrina berteroana responds
to regular pollardiig with a great volume of large leafed shoots in only 3
weeks to 1 month. These shoots are often used for rabbit feed and are also
palatable to livestock. Goats are particularly eager to eat these shoots.
Production: At CATIE (elevation 610 m, mean annual temperature
22.3"C and annual rainfall 2700 mm), a 100 m fence of E. berteroana
after pruning and carefully measurements showed the following produc-
tion (Table 3). The posts had an average spacing of 0.6 m and the lopped
branches were 4,6, and 8 months old after being pollarded.
A member of the Liliaceae, Yucca elephantipes Regel, with long stiff
dagger shaped leaves is notable for its large paniculate, whitecreamish,
bell-shaped flowers.These are sought out as a delicious vegetable which is
82 JOURNAL OF SUSTAINABLEAGRICULTURE
. - - - . .. - .
Table 3: Biomass production (dry weight) of one kilometer of fence
of Erythrina berteroana pruned every 4, 6 and 8 months.
Average distance between posts: 0.6 m.
(kg d. w. /prunning)
cooked with eggs. The stigma is usually removed to avoid a bitter taste.
Flowers are sold at US$0.50-1.00 her cluster in Costa Rican markets. Each
stem produces one cluster per year. It is rapidly propagated from cuttings
in most types of soil under dry and wet climatic conditions from sea level
to about 1500 m altitude. The plant is also used to stabilize terrain along
roads, particularly when there are landslides. Anacardium occidentale, the
cashew, will grow on very degraded soils in relatively dry climates. Psi-
dium guajava, the well known guava fruit, is used for jellies, canned for
juice or eaten raw, although usually the latter is infested with larvae when
ripe. This tree also does well on poor and compacted soils. The fruits like
those of various Ficus spp. are relished by cattle ,and other domestic
animals and in fact the seeds are spread through excrement. The wood is
frequently used for fuel.
Bursera simaruba exudes on oleo-resin called elemis. This aromatic
essential oil played a role in commerce in the past and provided a sticky
base for varnishes, printers, inks and in some ointments where stimulant
and antiseptic qualities were required. Home use is made of the gum as a
cement for mending crockery and as a crude varnish. The thick colored
sap exuded from wounds in Spondias purpurea is used in medicine.
CONCLUSIONS
The maintenance and other management practices of living fences vary
with the species, the ecological conditions, the local markets and the
cultural backgrounds.
Living fence posts are of relatively low cost, in comparison with normal
wooden "dead" posts, particularly when live fences from which cuttings
can be obtained are found nearby. They are also more durable, and rows of
live fence posts well over 50 years have been recorded.
Research, Reviews, Practices and Technology 83
But there are also drawbacks when using live fence posts in comparison
with non-living wooden, metallic or cement fence posts. They require
much more care for their establishment and maintenance, such as careful
handling of the large cuttingsprior to planting, and their possible replace-
ment if they do not take root. Periodic pruning is required and often
constitutes a main management problem, as can be witnessed by many
abandoned live fences. Some of the advantages and drawbacks concerning
live fences were previously presented (Budowski, 1982).
Live fences may become weeds and grow tall when they are not main-
tained. They can, and sometimes do, harbor pests. They may also be
considered as obstacles to the use of machinery and even for small air-
planes that spray pesticides. Under certain conditions, they compete for
water, nutrients, and light with adjacent crops, although over limited sur-
faces. Some of the fruits, and even the fofiage of some species, can be
toxic if ingested by livestock. And once planted live fences are difficult to
move or to eradicate because of their rooting systems. They are not easily
relocated, such as with electric fences or even dead fence posts.
Despite the important role of live fence posts in the rural landscape and
the vast amount of empirical knowledge accumulated, information is
scarce. More needs to be known about the cultural, edaphic and climatic
requirements of the many species used and the anatomical and physiologi-
cal basis of propagation from large cuttings and the various factors affect-
ing their regenerationand rooting.
Certain superior varieties among fence post species seem to exist, in
terms of growth habits, vigor, desirable architecture, sprouting behavior,
absence of spines ( i what would be normally spiny varieties), quality of
edible fruits and flowers produced, protein content of foliage and resis-
tance against drought or swampy conditions. Local people speak of vari-
eties and races, but these are tittle known since the information has not yet
been validated, much less quantified. The same may be said about the
ability of living fence posts to contribute to soil stabilization, to serve as
windbreaks or to supply sources of animal feed or forage. For the latter,
additional data about their protein content, digestibility, drying require-
ments or their pelleting capability is needed.
There is no doubt that a great potential exists for the development of
improved strains through the application of known selection and varietal
improvement techniques. Much potential exists for commercialization of
some of the multitude of derived products obtained from live fences and
the subsequent generation of additional income, particularly for the small
farmers.
Finally, the living fence is indeed a very promising practice to be
84 JOURNAL OF SUSTAINABLE AGRICULTURE
BIBLIOGRAPHY
Alavez Lopez, S. 1983. Estudio p r e l i i a r de 10s c ~ c o vivos
s en la ganaderia de
Teapa, Tabasco, (manejo de Arboles en 10s potreros, una tdcnica silvopastoril).
Tesis 1ng.A~.Chapingo, Mexico, Universidad Aut6noma de Chapingo. 77 p.
Allen, P.H. 1977. The rain forest of Golfo Dulce. Stanford, California, Stanford
University Press. 417 p.
Baggio, A.J. 1982. Establecimiento, manejo y utilizacidn del sistema agroforestal
cercos vivos de Gliricidia sepiurn (Jacq.) Steud., en Costa Rica. Tesis Mag.Sc.
Turrialba, Costa Rica, UCRCARIE. 91 p.
Beer, J. 1987. Experiences with fence lime fodder trees in Costa Rica and Nicara-
g u a In Advances in Agroforeshy Research Proceedings, Edited by J. Beer,
H.W. Fassbender and J. Heuveldop. CATIE. Tunialba, Costa Rica (Inf. Tec.
no. 117)
Beliard, C.A. 1983. Resultados preliminares de la producci6n de biomasa en
cercos vivos de Gliricidia sepium bajo dos frecuencias de poda en la regi6n de
La Palmera, San Carlos, Costa Rica Trabajo presentado en el curso Corto
sobre Metodologias de Investigaci6n Agoforestal en el Tr6pico Hiunedo
UNU/CATIE-IICA TROPICOS-CONIF, Cali, Colombia, 26 de noviembre a 7
de diciembre de 1983. Tunialba, Costa Rica, CATIE. 12 p.
Beliard, C.A. 1984. Producci6n de biomasa de Gliricidia Sepium (Jacq) Stend, en
cercas v i v a bajo tres frecuencias de podas (3,6 y 9 meses). M.Sc. Thesis.
CATIE-UCR, Tunialba, Costa Rica, 97 p.
Bond W.E.T. 1944. Hedge plants in Northern Nigeria Tropical Agriculture (Trini-
dad) 21(12):228-230.
Budowski, G. 1977. Ago-forestry in the humid tropics, a program of work.
Tunialba, Costa Rica, CATIE, Departamento de Ciencias Forestales. 24 p.
(report mimeo).
Budowski, G. 1982. The socio-economic effects of forest management on lives of
people Living in the area: the case of Central American and some Caribbean
countries. In Socio-economic effects and constraints in tropical forest manage-
ment Edited by E.G. Hallsworth. New York, John Wiley, 1982. pp. 87-102.
Budowski, G. 1987. Living fences in tropical America: a widespread agroforesby
practice. In Agoforeshy: realities, possibilities and potentials. Edited by H.L.
Gholz. Martinus NijhoR Publishers, Dordrecht, The Netherlands. pp.169-178.
Budowski G., D.C.L. Kass and R.O. Russo. 1984. Leguminous trees for shade.
Pesquisa Agropecuaria Brasileira 19(s/n):205-222.
Research, Reviews, Practices and Technology 85
Budowski, G., R.O. Russo y E. Mora. 1985. Productividadde una cerca viva de
Erythrina berteroana Urban en Turrialba, Costa Rica. Turrialba 35(1):83-86.
Burgos, J.A. 1952. Postes vivos para cercos. T i g o Marla, P d . Estaci6n Exper-
imental Agrkola 6 p. (Circular no. 39).
Centro Agronornico Tropical de Investigaci6ny Enseiianza 1986. Bibliography
on the genus Erythrina. CATIE, Renewable Natural Resources Department,
Turrialba, Costa Rica. 182 p.
Combe, J. y G. Budowski. 1979. Clasificacidnde las tknicas agroforestales: una
revisidn de litexatura. In G. de las Salas e d Taller de Sistemas Agoforestales
en Am6rica Latina. W a l b a , Costa Rica, CATIE, pp. 1748.
Cook, 0.E 1901. Shade in coffee culture. Washington, D.C., U.S. Department of
Agriculture, Division of Botany. 79 p.
Crane, J.C. 1945. Living fence posts in Cuba Agriculture in the Americas
5(2):34-38.
Duke, J.A. 1972. Isthmian ethnobotanical dictionary. Fulton, Maryland, Duke,
96 p.
Espinoza, J.E. 1984. Caracterizaci6n nubitiva de la fracci6n nitrogaadit del
fomaje de madero negro Gliricidia xepium y por6 Erythrina poeppigiana.
M.Sc. Thesis CATLE-UCR W a l b a , Costa Rica. 90 p.
Flinta, CM. 1960. Rdcticas de plantaci6n forestal en Adrica Latina. Roma,
FAO. 449 p.
Fonseca, M.T. 1968. El por6. Revista de Agriculma (Costa Rica) 40(6-7):
102-112.
Garcia W J . 1974. Emaizamiento de estacas de seis especies forestales con tres
niveles de 6cido indol-butirico.Tesis Mag.Sc., Twilaba, Costa Rica IICA 41 p.
Halliday, J. and P.L. Nakao. 1982. The symbiotic affities of woody species
under consideration as nitrogen-fixing trees. NiffAL Project, University of
Hawaii. 85 p.
Hardwood, R.R. 1979. Small fann development. Understanding and improving
farming systems in the humid tropics, Boulder, Colorado, Westview Press.
160 p.
Hartmann, H.T. and D.E. Kester. 1983. Plant Propagation: principles and practi-
ces. 4ed Englewood Cliffs, New Jersey, Prentice-Hall.727 p.
.Holdridge,L.R. 1966. The life zone system. Adansonia 6(2): 199-203.
Holdridge, L.R. 1970. Manual dentrol6gico para 1000 species arb6reas en la
Repdblica de Panam4 Rograma de las Naciones Unidas para el Desarrollo y
FAO. Panarni 325 p.
Holdridge, L.R. y L. Poveda. 1975. kboles de Costa Rica v.1. San Jose, Costa
Rica, Centro Cientffico Tropical. 546 p.
Howes, F.N. 1946. Fence and barrier plants in waxm climates. Kew Bull of
Miscellaneous Inf. 251-87.
Krukoff, B.A. 1939. The American species of Erythn'na. Brittonia 3:205-337.
Little, E.L., Jr. Common fuelwood crops: A handbook for their identification.
Morgantown, West Viginia, w.d, 354 p.
86 JOURNAL OF SUSTAINABLE AGRlCULTURE
Logsdon, G. 1978. Living hedge fences, shrub and trees varieties, planting and
pruning. Org. Gard Farming 25(5):78-84.
Lozano J., O.R. 1962. Postes vivos para cercos. Tesis Mag.&. Turrialba, Costa
Rica, IICA. 77 p.
Martin, F.W. y R.M. Ruberte. 1980. Techniques and plants for the tropical subsis-
tence fann. Mayaguez, Puerto Rico, U.S. Department of Agriculture. 56 p.
Martinez, M. 1959. Plantas dtiles de la flora mexicana. Mexico, D.F., Botas 621 p.
Mintz, S.W. 1962. Living fences in the Fond des-Neges region, Haiti. Econoniic
Botany 16:lOl-105.
Morton, J.F. 1981. Atlas of medicinal plants of Middle America. Illinois, Thomas.
1420 p.
Nobhan, G.P. and T.E. Sheridan. 1977. Living fence rows of the Rio San Miguel,
Sonma, M6xico: traditional technology for floodplain management. Human
Ecology S(2): 97-111.
National Academy of Sciences. 1979. Tropical legumes: resource for the future.
Washington, D.C., N.A.S., 326 p.
National Academy of Sciences. 1980, 1983. Firewood crops. Shrub and tree
species for energy production. V.l, V.2. Washington, D.C., N.A.S. 237 p. and
89 p.
Nuiiez MelBndez, E. 1978. Plantas medicinales de Costa Rica y su folclore. San
Jose, Costa Rica, Editorial Universidad de Costa Rica. 318 p.
Otarola T., A. y Martinez H., H.A. 1985. Manejo y producci6n de cercas v i v a de
Gliricidia sepium en el Nordeste de Honduras. Paper presented in Second
Meeting of I.U.F.R.0 Group S1.07.07-Agroforestry, CATIE, Tumalba, Costa
Rica, June 24-28,1985.
Picado, W, y R. Salazar. 1984. Produccidn de biomasa y leiia en cercas vivas de
Gliricidia sepium (Jacg.) Steud de dos aiios de edad en Costa Rica. Silvoen-
ergia (Costa Rica) 1:l-4.
Pittier, H. 1957. Ensayo sobre plantas usuales de Costa Rica. Zed rev.; Editorial
Universitaria. 186 p. (Serie Ciencias Naturales no. 2).
Russo, R.O. 1984. Studies on Erythrina poeppigiana (Walpers) O.F. Cook, a
multipqose tree in Costa Rican farms. Ph.D. Dissertation Southeastern UN-
versity, New Orleans, Louisiana, U.S.A., 1984. 148 p.
Sanchez, G.A. and R.O. Russo. 1985. Biomass production of Etythrina spp.
(Papilionaceae) in the humid tropics. Paper presented to the Conference "Re-
search in Foreshy for Energy." Copenhague, Denmark, October 28-November
1,1985.
Sanchez, G.A. and L. Payne. 1987. Survey of the cultural practices and uses of
Gliricidia sepium by farmers in Costa Rica In Gliricidia sepium (Jacq.) Walp.:
Management and Improvement. Edited by D. Withington, N. Glover and J.L.
Brewbaker. Nitrogen Fixing Tree Association, Waimanalo, HI. pp. 8-13.
Sauer, J.D. 1979. Living fences in Costa Rican agriculture. Turrialba (Costa Rica)
29(4):255-26 1.
Schroeder, C.A. 1976. Living fence posts in Central America. Los Angeles,
Reseatch, Reviews, Practices and Technology 87
RECEIVED: 5/14/91
REVISED: 6/5/92
ACCEPTED: 6119/92