Vous êtes sur la page 1sur 3

tacf /, ka6/c-

/ldaA/ftU//Ad4
' a( a/rtd/44 rtMzce
/ '/ /APP//
/-l,t md,r a- /2t4+t
TA:ZaraTabi(ztabi@ucsd.edu) BICD120:
Dr.Schmidt
AIM:ztabi06 Week2
Details
TheNittv-Grittv
Photosvnthesis:
+ 12HzO+
6COz + 18ATP
12NADPH + 60z+ 12NADP-+
) CoHrzOo 1BADP+ 6 HzO

TheLiqhtReactions: ofATPandNADPH
Production fromlightenergy!
- lmportant PigmentsinPhotosynthesis
( . Primary pigment;
photosynthetic a) located
Chlorophyll inthereaction ofeachphotosystem
center
^rrrrmrrtira
"' .
S,,|tfi:l | Secondary/Acessory Pigments;Helpwidenspectrum byplant) located
oflightthatcanbeabsorbed inthe
'il[n[T^t, I UghtHarvestingcomplex(LHC) aL &//Hna catdzlr'zx
+l-tlLu( \ b,c,etc.
- Chlorophyll - Carotenoids
rt' .
Ynld,npnoYl Beabletointerpret graphs andAction
oftheAbsorption Spectral
fficnrt' whatwas
- Understand from
concluded T,W.Engleman's experiment
= SizeofLHC
-Photosynthetic'unit
. SizeofLHChelps preventPhotooxidation theamount
bycontrolling oflightabsorbed
- Excesslightthecause ofphotooxidation
which forms fee radicals
cell-damaging
- Carotenoidsabletogather toprevent
freeradicals damage
. Growing inshade ) Large sizeofLHCtoabsorb morelightenergy) higherchlorophyll
B/ARatio
. Growing insun) Small sizeofLHCtoregulatelightintake
) lower chlorophyll
B/Aratio

- Know
thename ofeachprotein
andlocation complex:
XM f 26o +o t 4 \oz +.o /,
- Peffictdes offie-cnnn ph o\sflm)vtls ;
o t1^4- pvatervl @wptexes
Pt-wnl'\ e- fra,r,rsporl CA ov€
"t
- &+l.r.cfl\^e PvbteiYl rch^af!\s oX'l?\ZiiD Ccayr'tgarn e-)
' Tl4oueb<W ,t, =) R.EBTcAD (Xain e- bu1 rtowM)t 't-" lrr"J t,
. Jhosc A-Ef€/t-1f :) oxtO\ZOD

- cyctic photophosphorytation
Vs,Noncyctic thcse*fi:?*f, it"
. InNon-byclic
phoiophosphorylation: )
rl {rowr - useofBoTH ll
Photosystem and| ./ fr OFi *fn
-y.'?2'
olN ArP
}-f*rDd<-
{litrnfi
of 2H2Oproduces
4 e-,102 2 NADPH ' we.rced.a
\ uoo' " \ - Splitting ,.'
,r"^1_'q,):3il,1iffiili1'i.'lljx:1'J:ryi photophosphoryra
produced
via ,

s,1${0rnfr
ffii::trifr3J:4TP
\
. lnCyclic
photophosphorylation,
noNADP+ isavailable
tobereduced,
soe-areshuttled
backto
topumpmoreH+) MoreATPProduced
Plastoquinone
- | ) NoH2Oissplit€
onlyPhotosystem
Uses ,o oz tgaa.tcd

4 rwn'cV&d fbryMuag//nhon cet4 oe'r /4 /hz 'sa'////


x ealfie /?ar 'Mh feo
ull, n nlrw?b/hil/ -) r,ot r&/vr'fu h dltau 7/aztR
'1at//6fe
r/pq/a///co V ?U" pr/h'/r6Qtry//ana7u' tfr /a/urt ry//-
,/ TheCalvinCvcle/Dark Reactions/C3
Photosynthesis foodsource
Producing fromCO2Fixation
- DiscoveredbyMelvin Calvin 7
fixedintoa 3-Cmolecule
- C)zinitially -/uhf/v.r/ frs ahz/ Pelz
(3-phosphoglycerate)
-ex?e^vvlenIed *>/ rMtoAriive e +o &tAlMvW flnat 6Ye i> 1\a.r-
tnl,[tCio vVwe-64l\Q Coz t ,-[Ved, tvt-
- $o need t> bvrouJ aftuc/rursg cph ,\w-Le.- wv>epl,o\eL , bot
bnoo uJna+ art-c.- et\d .ft\t/ ftdl/L
Y W f6%,r. U,(A+eOl)
ab w)L aa .l\etr qluAfi%

VvlPnfMt f\ Evt ot-t>


t A?@'altA Clqd.i"cPhopPhosphovgtrd-ron
i3
r/ "C-Hg
- '
- Need3 ATPand2 NADPH perco2 Fixed "#'X.,t
- TheProblem withRubisco
) hasanaffinityforBOTH CO2and02
' flow02]and[HighCO21= Rubisco binds toCO2=CO2Fixation
7 ' [high02]and [LowCO2]= Rubisco binds to02 = Photorespiration
= NetCO2Loss!
L th,. pratvra[tsu4ts sr>tredbg a lrn sonleplarats)
C4PhotosvnthesisAnother methodofcarbon fixfiion,discovered
byscientists
Hatch andSlack(1965)
- WhyCalled C4?) COzFixed endsupina 4-Cmolecule (Oxaloacetate)
insteadof3-C!
- C4andC3photosynthesis arespatially
separated-Different cells
' /// C(p&4tfr A1z7apifu Cl * C6lnaas/tt**tts
4 /ylfrnrlltr'oAdl
- fnilk / lzz CU7z>a*' a,a ,,
au rb C€

-Whydoitwhenit usesupmoreenergy phosphates,


(intheformofhighenergy whenATP) AMP)?
Makes a MORE EFFICIENT PROCESS:
1)PEPCarboxylase hasa higher forCOz
affinity incomparison
toRubisco
2)Createanenvironmentof +t/7 &tat//ee?ZlV a&s
high[CO2]tofavorCO2fixationoverphotorespiration
3i Arrangementofcells(C4orkrinzAnatomy) helps CO2thatmayhavebeenlostduring/ut/t/, ,h4rsoo
recapture
photorespiration b/.ntrltn 6ua op2 ,fl ffu,4 /u a2/77't/
/ Ae, r6u &,YZ'o
- BecauseCO2lossisrcduced, canclose
stomatamore often) lessH2Ol'oss!
. ldealforplants ofhighlightandtemperature,
thatliveinareas rateofphotorespiration
where increases
ex.TropicalGrasses-Maize,
Sugar Cane, Millet,
etc.
Photosynthesis-
AcidMetabolisd
CAMGrassulacean ofC4photosynthesis
version
Analternate
aretemporally
- C4andC3Reactions andNight
separated-Day
- nZ/nnV At nrb AAA, /af q/e'/Vuo/2t27 m6/dz ru aZ
%
- Oe-atrrrL2a/ /uAbPl2 patd rnstdc ,/* *z-'7a"ffi7

- Becausehigh[] ofmalate (malicacid) inthevacuole


stored pHwillusually
atnight, belower
- Whydiditevolve?
' Helpplants withliftlerainfall
thatliveindrylocations
- Keepstomata thedaytime
during
closed topreventwaterloss) Canstillperform
C3photosynthesis
- Openstomata whentemperature
atnight totakeinCO2toperform
iscooler C4photosynthesis
. ex.Cacti,succulents, pinepple

PrimarvProductivity,
GlobalWarminq-Questionstothinkabout:
productivity?
forprimary
- Definition
- Whattypes ofplants
(C3,C4,CAM, atfixing
etc.)arethemostefficient Least
CO2? efficient?
ecosystem
- Whatcontinental playsanimportantroleinfixing peryear?
themostcarbon
- HowmanyPounds ofCO2 are for
released gallon
every ofgasused?

(Referr given
tothehandout lecture!)
on10i08109