Wednesday,August24,2011

UniversityofColoradoSchoolofMedicine
GraduatePrograminCellBiology,StemCellsandDevelopment(CSD)
StudentHandbook
20112012AcademicYear

I.

Page

Mission2

II.GraduateSchoolAdministration..........................3
III.

ProgramComponents
Year14
ThePreliminaryExam................................7
Year28
TheComprehensiveExam............................9
Year3andbeyond11
ThePhDThesis.......................................12

IV.GraduateSchoolStandards.......................................15
V.ObligationsandRecordKeeping.................................16
VI.OtherProgramEvents............................................17
AdvancetopicsandElectives...................................19
CurrentStudentsoftheProgram.................................20
ProgramFaculty..............................................21
Committeemembership........................................22
DegreesConferred............................................23

UniversityofColoradoSchoolofMedicine
GraduatePrograminCellBiology,StemCellsandDevelopment(CSD)

Director:LindaBarlow,Ph.D.
ProgramAdministrator:JenniferThurston
Mailingaddress:
UniversityofColoradoSchoolofMedicine
GraduatePrograminCellBiology,StemCellsandDevelopment
MailStop8108
12801East17thAvenue
Aurora,CO80045
http:/www.ucdenver.edu/csd
Phone:303-724-5902
FAX: 303-724-3420

Thishandbook,doesnotconstituteacontractwiththeUniversityofColoradoSchoolof
MedicineGraduateSchoolortheCellBiology,StemCellsandDevelopmentGraduateProgram,
eitherexpressedorimplied.TheCellBiology,StemCellsandDevelopmentGraduateProgram
reservestherightatanytimetochange,delete,oraddtoanyoftheprovisionsatitssole
discretion.Furthermore,theprovisionsofthisdocumentaredesignedbytheCellBiology,Stem
CellsandDevelopmentProgramtoserveasfirmguidelinesratherthanabsoluterules,and
exceptionsmaybemadeonthebasisofextenuatingcircumstances.

I.

Mission

TheprimarygoaloftheCellBiology,StemCellsandDevelopment(CSD)Programistotrain
talentedscientistsincellanddevelopmentalbiology.TheProgramstrivestoattractoutstanding
studentswiththehighestpotential,andtoprovidethemwithqualitytrainingthatstimulates
independentandcreativescientificthinking.Ultimately,helpingstudentstodeveloptheirfullpotential
inbecomingindependentinvestigatorsandleadersinbiologicalscience.
TheProgramsemphasisisonthedefinitionandresolutionofbiologicalproblemsratherthan
theapplicationoftechnologies.Thematically,theprogramisfocusedoncell,stemcellsand
developmentalbiologyandoffersawiderangeofresearchopportunities.Thenatureofthisprogram
willbestservethosestudentswhoareinterestedindevelopingindependentresearchcareersandwho
wishtopursueproblemsinbiomedicalsciencefromaninterdisciplinaryperspective.
Aftertheinitialperiodofcoursework,studentschoosetheirspecialtyfieldsfromadiverselist
oftopics,andproceedwithresearchuntilthegenerationanddefenseofathesisleadstotheawardofa
Ph.D.inCellandDevelopmentalBiology.

II.

GraduateSchoolAdministration
TheGraduatePrograminCellBiology,StemCellsandDevelopment(CSD)ispartofthe
GraduateSchooloftheUniversityofColorado,whichisanequalopportunityinstitution.Thegraduate
programsattheAnschutzMedicalCampusarepartoftheGraduateSchool,whichisunderthe
leadershipofBarryShur,PhD,Dean;andFranOsterberg,AssistantDean.
TheUniversityofColoradoDenverGraduateSchoolCampuspublishestheGraduateStudent
Handbook,whichincludesgeneralinformationandrulesconcerninggraduatestudents,aswellas
specificinformationonHonorCodeandGrievanceProcedures.Thisinformationappliestostudentsin
allprograms.ThepurposeofthishandbookistorelayadditionalinformationspecifictotheCSD
program.
A.
StudentSupport.Atpresent,studentsacceptedinthePh.D.programareprovidedfulltuition,
healthinsurance,andastipendof$26,000peryearforlivingexpenses(fortheacademicyear2011
2012).Continuedsupportiscontingentuponsatisfactoryacademicandresearchperformancebythe
student.Whenastudententersathesislab,thethesismentorassumescompleteresponsibilityforthe
studentsstipend,tuition,fees,andassociatedresearchcosts.Inordertoqualifyforinstatetuitionfor
thefollowingyear,alloutofstatestudentsmustestablishColoradoresidencybythesummerof
thefirstyear.
B.
StudentAdvising.Duringthefirstyear,CSDstudentswillmeetwithmembersoftheGraduate
AdvisoryCommittee(GAC)onarotatingbasistodiscussthestudentsprogressintheCSDProgram
andanyquestionsthatmaycomeup.Studentswillbeexpectedandencouragedtoseekadvicefrom
the GAC, Director, and/or other CSD faculty and student members prior to lab rotations,
ComprehensiveExamination,andanyothersituationrequiringfacultyconsultation.

III.

ProgramComponents

1STYEARSTUDENTS
COURSEWORK,LABORATORYROTATIONS,ANDPRELIMINARYEXAMINATION

FallSemesterRequiredRegistration
BiomedicalSciencesCourseCore
IDPT781112131415
10units
CourseDirector:Drs.M.Churchill,R.Davis,R.Evans,A.Bradford,K.Artinger,
Thissectionofthecoursecoversstructural,cellularandmolecularbiology.
EthicsinResearch
PHCL7605
1unit
CourseDirector:Dr.JimSikela
Courseisdesignedtointroduceissuesaroundethicsofresearch,publication,and
reviewingofmanuscriptsandgrants.
ResearchinCSDV(LabRotations)
CSDV7650(OV1&OV2)1uniteach
(registerforbothsectionsOV1andOV2)
CoordinatedbytheGACChair,Dr.TomEvans.
Studentswillperformresearchinthelaboratoryofoneofthemembersoftheprogram.
Therotationwillbefollowedbyanoralpresentation.
Cells,Development,andCancerSeminar
Noregistrationrequired
0units
CourseDirector:SeminarCommittee
Seminarseriesdesignedtopresentrecentimportantfindingsincellanddevelopmentalbiology
research.DifferenttopicsarepresentedweeklybyCSDTrainingProgramfaculty,studentsand
visitingfaculty.Attendanceisrequired.Seminarscheduleswillbedistributedbythe
ProgramAdministratoreachfallsemesterandarealsoavailableontheprogramswebsite.

SpringSemesterRequiredRegistration
StemCellsandDevelopment
CSDV7605
4units
CourseDirector:Dr.BruceAppel
Coursewillcovertopicsinthedevelopmentoftheorganism,suchascellbirth,migration,
differentiation,anddeath,inavarietyofdevelopmentalsystems.Partofthecourseisdevoted
tothediscussionoforiginalliteratureinthefield.
1Elective
(seep.20foralistofcourses)
CourseDirector:Variespersemester
Coursewillfocusonspecifictopicseachsemester.
CSDV7650(OV1)

ResearchinCSDV(LabRotation)
(for3rdlabrotation)
4

variable

1unit

CoordinatedbytheGACChair,Dr.TomEvans.
Studentswillperformresearchinthelaboratoryofoneofthemembersoftheprogram.The
rotationwillbefollowedbyanoralpresentation.
Cells,Development,andCancerSeminar
Noregistrationrequired
0units
CourseDirector:SeminarCommittee
Seminarseriesdesignedtopresentrecentimportantfindingsincellanddevelopmentalbiology
research.DifferenttopicsarepresentedweeklybyCSDTrainingProgramfaculty,studentsand
visitingfaculty.Attendanceisrequired.Seminarscheduleswillbedistributedbythe
ProgramAdministratoreachfallsemesterandarealsoavailableontheprogramswebsite.

SummerSemester
JoinaresearchlabonorbeforeJune15andstartonthesisproject.RegistrationisNOT
required(thispertainsto1styearstudentsonly).

A.
LaboratoryRotationsintheFirstYear.Rotationsserveseveralimportantpurposes.First,
theyenablethestudenttoexploreandcompareseveralareasofcellanddevelopmentalbiology
researchandaidinthechoiceofamentorandprojectforthesiswork.Second,rotationseminars
provideintensetraininginthecraftandartofpublicpresentation,anessentialaspectoffuturecareer
success.Third,theyallowprogramfacultytoevaluatethemotivationandintellectualpreparednessof
studentstoundertakeindependentresearch.Studentsshoulddiscusstheirinterestswithseveral
potentialfacultymentors,severalweeksormorebeforethestartoftherotation.
ROTATIONSCHEDULEFOR20112012:
Fall1st:August29November18,2011
Fall2nd:November21February24,2012
Spring:February27May18,2012
B.
NumberofRotations.Studentsmustperform3rotationsbeforethestartoftheirsecond
year.Studentsshouldstarttheirfirstrotationinthefallsemester.Studentsmustcomplete3rotations
in3separatelaboratoriesinordertoadvancetotheirsecondyear.Registerforthefirst2rotations
(Sections1&2OV1&OV2)inthefall;registerforyour3rdrotation(Section1OV1)inthespring.
OfficialdatesforrotationsaresetbytheGraduateSchool.Optionally,studentsmaystartarotation
duringthesummerbeforeformalentryintotheProgram.MedicalScientistTrainingProgram(MSTP)
studentsmustcompletetworotations(duringthesummersofthefirstandsecondyearofMedical
School).
C.
RotationExpectations.Forprofessionalsintraining,itisnotappropriatetorequirea
minimumnumberofhoursforrotationwork.Strongselfmotivationisanabsolutelyessential
characteristicforanindependentscientist,andweexpectourstudentstodemonstratethisquality
throughouttheirtraining.Inthisregard,studentsshouldexpecttobeinthelabbeyondthenormal
5

workinghours,i.e.evenings,weekends,andpossiblyovervacationdaysduringtheterm.This
commitmentoftimeisespeciallyimportantwhenlong,complexexperimentsarebeingdone.Amajor
partofthementorsrotationalassessment(aswellashis/herwillingnesstoacceptastudent)willbe
basedonthedegreeandqualityoflabeffort.Studentsshouldalwaysdiscusstimeoffand/orvacation
dayswiththeirlabmentorinadvance,bothintheirlabrotationsandoncetheyenterathesislab.
Ashortwrittenevaluationofthestudentsrotationwillbeprovidedbythefacultymentor.
Studentsarerequiredtogiveanoralpresentationoftheirrotationprogressaswellassubmitawritten
reportoflaboratorywork(12pages;includinghypothesis,necessarybackground,experiments
preformed,resultsobtainedandconclusion).Aftercompletingtheserequirements,rotationgradeswill
beassignedbythefirstyearadvisorinconsultationwiththerotationmentor,anddiscussedwiththe
student.
D.
RotationSeminar.Attheendoftherotationthestudentwillpresentaseminar.Thepurposeof
theseminaristoprovideintensetraininginthecraftandartofpublicpresentation,anessentialaspect
offuturecareersuccess.Eachseminarshouldbe15minutesinlength.Thestudentmustrehearsethe
seminarwithhisorherrotationmentorpriortothepublicpresentation.Theseminarisanessential
componentoftheresearchrotation.Studentsareexpectedtopresentawellorganized,clear,and
thoughtfulseminar.Studentsshouldconsiderthefollowingelementswhendesigningtheir
presentation(althoughtheorderneednotbestrictlyfollowed):
Introductionashortstatementofthequestionorproblemaddressedbytherotation,andthehypothesis
tobetested.
Background describe the significance of the question in broad terms for a diverse audience.
Describepreviousworkanditsrelationshiptotheproject.
Specific experimental aims what were the particular experimental goals proposed to test the
hypothesis?
MethodsandDesignexplainbrieflyanyunusualstrategiesortechniquesemployed.
Results.negativeandpositiveresultsshouldbereported
Conclusionsandfuturedirections whatcanyouconcludefromyourresults,andwhatwouldyou
pursueifyouremainedontheproject.
SuggestionsforEffectiveSeminars
1. Avoidreadingormemorizingyourpresentationwordforword.Wooden,canneddeliveriesare
dullandveryhardforaudiencestofollow.
2. Prepareandusesimple,effectivevisualaids.Rememberthateffectivecommunicationofdataand
ideasisyourgoal!Donotspendundueeffortandexpenseonfancymulticoloredslides(especially
fortext),ifcolorisnotrequiredtosimplifycomplexdataorconcepts.Coloredvisualstendto
requireadarkenedroomandareoftenmuchhardertoreadthanblackonwhitelinedrawingsor
letters.Keeptextverybriefanddonotreaddirectlyfromthescreen(audiencesaremuchfasterat
readingsilently!).
3. Usethemarkerboardwhenappropriate.Diagrammingoroutliningwhileyouaretalkingisa
highlyeffectivemeansofexplainingconceptsdifficulttodescribewiththespokenword.Useof
themarkerboardcanalsohelpanswerspontaneousquestionsfromtheaudience.
6

4. Consideraudiencequestionscarefully!Bothfacultyandstudentsareencouragedtoaskquestions
duringandafterrotationseminars.Afewofthesequestionsmaybeintendedtoprobeyour
understandingofyourresearchratherthanilluminateanareaofconfusion.Partofyourevaluation
willconcernyoureffectivenessinrespondingtoquestions.Thus,makesurethatyouunderstand
thequestionbeforeanswering.Repeatthequestionoraskforarephrasingifyouneedto.Second,
relaxandtakeamomentofsilenceifyoumustbeforeansweringtoformulateacoherentanswer.
Third,ifaftercontemplationyoudontknowtheanswer,dontbeafraidtosayso.Weallget
stumpedfromtimetotime!
E.
TransfertotheThesisLabatEndofFirstYear:Animportantaimoftherotationsisto
enablethestudenttoobtainathesismentor.Afterthecompletionofthethreerotationsforregular
graduatestudentsortworotationsforMSTPs,thestudentmustcometoamutualagreementwitha
facultymembertoactastheirthesismentor.ThechairoftheGACandtheProgramAdministrator
mustbenotifiedonthechoiceofmentoronorbeforeJune15thofthefirstyear.Officialtransferto
thethesislabtakesplaceonJuly1st.Underexceptionalcircumstancesandatthediscretionofthe
GAC,astudentmaybeallowedtoperformanadditionalrotationduringthesummerfollowingthefirst
academicyear,fortheexpresspurposeofenhancingthementorselectionprocess.
F.

PreliminaryExamattheEndoftheFirstYear:

1.ThegeneralformatofapreliminaryexaminationfortheCellBiology,StemCellsandDevelopment
GraduateProgramisawrittengrantproposalfollowedbyanoralexaminationbyapreliminary
examinationcommittee.
2.Thepreliminaryexaminationcommitteewillconsistoffivefacultymembers.Everyyear,following
thefirst2yearsafterinitiationofthispreliminaryexamformat,twocommitteememberswillbe
replacedwithnewfaculty.Eachmemberwillserveaminimumoftwoconsecutiveyears.The
committeewillalsoconsistoffacultyrepresentingdifferentaspectsoftheresearchwithinCSD,such
asDevelopment,CellBiologyandStemCellBiology.
3.Fourweeksbeforetheoralexamination,studentswillbeprovidedwithfiveresearchtopics;one
topicfromeachcommitteemember.Eachtopicwillberepresentedby23papersthathavebeen
selectedbythecommitteemembers.Eachstudentwillneedtopickonetopicforhis/herproposal.
Whilestudentscanselectthesametopic,obviously,studentsarenotallowedtoworktogetherontheir
proposals.Thetopiccannothaveasignificantoverlapwithstudentsresearchinterestsintheirfuture
lab,andwillhavetobeapprovedbythecommittee.
4.Eachstudentwillhaveonecommitteememberassignedasapreliminaryexaminationmentor.The
samecommitteememberwillalsoserveasachairduringexaminationofthisstudent.Themainroleof
thementorwillbetoserveasagotopersonforthestudentifs/he(thestudent)hasquestions
regardingthewrittenandoralportionsoftheexamination.Thementorcanadvisethestudent
regardingtheexpectationsofthewrittenandoralexaminations.Thementorcannot,however,be
directlyinvolvedineditingorrewritingthestudentsgrantproposal.Mentoralsocannotbedirectly
involvedinsuggesting/designingtheexperimentsorinterpretationsofpotentialoutcomesthatwillbe
describedintheproposal.
7

5.Studentswillcompletethewrittenproposalanddeliverittoeachmemberofthecommitteenolater
than1weekbeforethedateoftheoralexamination.Thisdeadlineisfirm.Theproposalistofollow
theNIHpredoctoralfellowshipformatandcanbenolongerthan6pages(excludingreferences).
6.Inadditiontothewrittenproposal,thestudentwillbeexaminedorallybythecommittee.The
examinationforeachstudentwilllastapproximatelyonehour,unlessthecommitteedecidesadditional
timeisneeded.
7.Theexamisdesignedtotesteachstudentsunderstandingofkeyconceptsandabilitytothink
throughexperimentaldesign,bothofwhichareimportantforresearchinbiomedicalsciences,witha
focusondevelopment,cellbiologyandstemcellbiology.Whilethemainfocusofthequestionswill
berelatedtothewrittenproposal,studentsshouldexpectquestionsoutsidetheimmediatescopeof
writtenproposal.Allquestions,however,willbelimitedtothematerialthatthestudentwasexposedto
duringcoursesandrotationsthattheyhadwithinthefirstyearofagraduateprogram.
8.StudentswillbegradedonaPassorFailbasis,asdecidedbyamajorityofthecommittee.Thefinal
gradingwillincorporateacombinedevaluationofthewrittenproposalaswellasoftheoral
examination.Bothaspectsoftheexaminationwillbeweightedequally.
9.Intheeventofafailedexam,thesituationwillbereferredtotheGraduateAdvisingCommittee
(GAC)forCSDforaremediationandexamretakeplantobecompletedbytheendofthesummer
semester.

2ndYearStudents

COURSEWORK,PRETHESISRESEARCHANDTHECOMPREHENSIVEEXAMINATION

FallandSpringSemestersRequiredRegistration
ResearchinCSDV
CSDV7650(OV3)
CourseDirector:Dr.LindaBarlow
LaboratoryresearchwithCSDTrainingProgramfaculty.

15units+

1ElectiveorAdvancedTopicsCourse
(seep.19forlistofcourses)
variable+

CourseDirector:Variespersemester
Coursewillfocusonspecifictopicseachsemester.
Canbetakenineitherthefallorthespring(afterthefirstyear,oneelectiveoradvancetopic
classisrequiredperacademicyear.
+

)thefallandthespringsemestersmusteachtotal5units

NOTE:

Eachfallsemester,allstudentsmustcompletetheformsforinsuranceplanselection.
ContacttheStudentInsuranceCoordinator,3033150800withquestions.Theseforms
mustbecompletedwhetherornotyouplantoparticipateinaStudentHealthInsurance
Plan.

SummerSemesterRequiredRegistration
DoctoralThesis
CSDV8990
1unit
st
Allstudents(except1 year,whohavedifferentrequirements)mustberegistered
duringthesummermonthstobeconsideredfulltimeandtoavoidhavingretirement
takenoutoftheirpaychecks.

Updatetalks:Beginninginthesecondyear,eachstudentisrequiredtogiveanannualupdate
presentationtotheprogram.ThefirstupdateshouldbebeforeMay31ofthe2ndyear,andshouldbe
scheduledwiththeprogramadministratoratleastonemonthinadvance,althoughthepresentationcan
bescheduledatanytimeduringtheyear,i.e.,earlierisbetter.Wealsorecommendthatyouforma
committeeandhaveoneprecomprehensiveexammeetingatthetimeofyourfirstupdateseminar.
Thus,toscheduleacommitteemeeting,youwillneedtobesureallyourmemberscanattend,and
coordinatingfacultyschedulescanbechallenging.Again,wewanttoemphasizethatarrangingthis
updateisyourresponsibility,andurgeyoutomakeplanswithyourcommittee,andscheduleyour
presentationwiththeprogramadministratorwellinadvance.

ComprehensiveExam.AtthebeginningofthesecondyearofstudyCSDgraduatestudentswillbegin
preparingfortheComprehensiveExam.Itishighlyrecommendedthatthestudentcarefullyreadthe
GraduateStudentHandbookonComprehensiveExaminationpoliciesanddeadlines,andpickupa
packetofinstructionsandformsfromtheGraduateSchoolwellaheadoftheplannedexaminationso
allrequiredpaperworkcanbecompletedontime.Completedpaperworkmustbesubmittedtothe
GraduateSchoolnolaterthantwoweekspriortotheexaminationdate.Note:Astudentmustbe
registeredatthetimehe/shetakestheComprehensiveExamination.
StudentsmusttaketheGraduateSchoolComprehensiveExaminationforadmissiontocandidacyfor
theCSDPh.D.bytheendofthesummeroftheirsecondyearbutbeforeJanuary1stofthethird
year.AnydeviationtothisrequirementmusthaveapprovalfromthePI,GraduateAdvisory
Committee,andtheProgramDirector.TheComprehensiveExaminationCommitteeshallconsistofa
minimumoffiveGraduateFacultymembers.Atleastoneofthemembersmustbeoutsidethe
Programscoretrainingfaculty.Themajorityofthemembers,includingthechair,mustbefromthe
coretrainingfacultyoftheCSDProgram.StudentsshouldcontactmembersoftheGraduateFaculty
whomtheywishtobeontheircommittee,inconsultationwiththeDirectoroftheProgramandtheir
thesisadvisor.Studentsshouldinformthecommitteemembersoftheirbackground,thetopicoftheir
thesisresearchandtheirpreliminaryresults.Inaddition,thestudentarrangesthetimeandlocationof
theexam,andinformsthemembersofthecommitteethattheexaminationrequiresthreehours.The
studentshouldprovidetheGACwiththenamesofthecommitteemembers.
Theexaminationwillhaveasitsfocusathesisresearchproposalwrittenbythestudentusingthe
formatofaNIHpredoctoralfellowship.Althoughpreliminarydatacollectedbythestudentare
helpful,itisnotessentialfortheproposal.Thewrittenproposalmustbedistributedtothe
ComprehensiveExamCommitteeatleasttwoweekspriortotheexamination.Thestudentmust
adequatelydemonstratethescientificknowledgeandabilitytodefendthisproposal,aswellas
satisfyingtheoverallrequirementsfortheexaminationassetforthbytheUCDAMCGraduate
StudentHandbook.Theexaminationwillconsistofa3045minuteseminarbythestudent,general
questionsfromtheaudienceandthendetailedquestionsfromtheThesisCommittee.Asstatedinthe
graduatestudenthandbook,thecomprehensiveexaminationwilltestyourmasteryofabroadfieldof
knowledge,notmerelytheformalcourseworkcompleted.Thestudentshouldconsultwithhisorher
committeememberspriortotheexamastothesubjectareaseachmemberexpectsthestudenttohave
mastered.
1.Writtenproposal
Listedbelowareguidelinesfortheproposal:
Title
ShortIntroductionandSpecificAims
1.0page
BackgroundandSignificance
2.0pages
PreliminaryStudies
2.5pages
(thissectionisnotessential)
(additionalresultscanbeplacedinanappendix)
ExperimentalDesign&Methods,
4.0pages(detailedforeachspecificaim)
ExpectedResults&Interpretationthereof
10

ImplementationTimeTable
Total

0.5pages
10.0pages

Literaturecitationsareadditionaltothe10pages.Fullreferenceswithtitlesarerequired.
Thewrittenproposalmustbegiventoallmembersofthecommitteeatleasttwoweeksbeforethe
comprehensiveexam.
2.TheExam
ThecandidateforthePhDshouldprepareatalkof3045minutesonhis/herthesisproposaland
preliminaryresults.Thetalkwillbeopentotheuniversitycommunity.Afterthetalk,questionsfrom
thoseinattendancewillberequested.Afterthequestionshavebeenaddressed,allbutthegraduate
facultyandtheComprehensiveExaminationCommitteewillberequestedtoleave.Therewillbetwo
typesofquestionsfromthecommittee:
1.Questionsonthewrittenandoralproposal:~1hr.
2.Generalknowledgequestionsprimarilyfromthecourseworkofthecandidate:~1hr.
Afterthequestionsthecandidatewillbeaskedtoleave,andthecommitteewilldiscusstheresultsof
thedifferentexamcomponentsandreacharecommendation.Thecommitteechairwillbringthe
recommendationsofthecommitteebacktothecandidateandfillouttheappropriateformsofthe
GraduateSchool.
3.PossibleResults
Pass(noconditions)
ConditionalPass(conditionsmustbedetailed)
Continuationofexamination(commentsmustbeincluded)
Fail(thestudentmustleavethegraduateprogram)
4.ApplicationtotheGraduateSchoolforAdmissiontoCandidacy
Applicationsmustbecompletednolaterthanthreeweeksbeforetheexam.Formsareavailablefrom
theGraduateSchoolandmustbeapprovedbytheProgramDirectorandreturnedtotheGraduate
SchoolOffice.ThedateoftheComprehensiveexaminationandthecompositionofthecommittee
mustberegisteredwiththeGraduateSchool.Thestudentmusthaveaminimumof30credithours
priortotheexamdate.
NOTE:

Afterpassingthecomprehensiveexam,allPhDstudentsarerequiredtoregisterfora
minimumoffive(5)hoursofDoctoralThesisCSDV8990(insteadofCSDV7650)
eachfallandeachspring.Failuretodosocanresultinthestudentbeingrequiredto
retakethecomprehensiveexam.Astudentmayregisterforupto10unitsofCSDV
8990inthesemestersbeforeandthesemesterinwhichthecomprehensiveexamis
takenandpassed.

11

3rdYearandAdvancedStudents

FallandSpringSemestersRequiredRegistration
1ElectiveorAdvancedTopicsCourse
(seep.19forlistofcourses)

CourseDirector:Variespersemester
Coursewillfocusonspecifictopicseachsemester.
Canbetakenineitherthefallorthespring(afterthefirstyear,oneelectiveor

variable

advancetopicclassisrequiredperacademicyear)

DoctoralThesis
CSDV8990
5+units
Allstudentsmustregisterforaminimumof5unitseachfallandeachspringafter
passingtheircomprehensiveexam.
Studentswillgenerateanoriginalbodyofresearchthatconstitutesasignificantcontribution
tothefieldofcellanddevelopmentalbiology.
SuitabilityofthesisresearchisjudgedbytheThesisCommittee.
StudentswriteaPhDthesisanddefendthedocumentatanoralexamination.

SummerSemesterRequiredRegistration
DoctoralThesis
CSDV8990
1unit
Allstudentsmustberegisteredduringthesummermonthstobeconsideredfulltime
andtoavoidhavingretirementtakenoutoftheirpaychecks.

NOTE:

Comprehensiveexam(seep.9)mustbetakenbyDecemberofYear03.

NOTE:

EachFallsemester,allstudentsmustcompletetheformsforinsuranceplanselection.
ContacttheStudentInsuranceCoordinator,3033150800withquestions.Theseforms
mustbecompletedwhetherornotyouplantoparticipateinaStudentHealthInsurance
Plan.

A.ContinuousRegistrationRequirement.
Studentsmustregistercontinuouslyfollowingsuccessfulcompletionofthecomprehensive
examination,i.e.,5creditsforfallandspringsemesterand1hourofthesisresearch(CSDV8990)
duringthesummer.Allstudentsmustregisterforboththefallandspringsemesters.Itisthestudents
responsibilitytoregisterforclassesinatimelymanneralllatefeesandfinancechargeswillbethe
responsibilityofthestudent.
B.ThesisResearch
Withinonemonthofthecompletionofrotations,eachstudentmustreachamutualagreementwitha
facultysponsorwhowillserveastheirthesisadvisor.Thechoiceofthesisadvisormustbeformally
approvedbytheGraduateAdvisoryCommittee.Thestudentandfacultymembertogetherplanathesis
project.Doctorallevelworkrequiresaclosecollaborationwithafacultymentor.Itistheresponsibility
12

ofthestudenttoestablishandmaintainthatrelationship.ContinuationinthePhDprogramdepends
uponthestudentidentifyingamutuallyagreeablethesislaboratory.Thethesisresearchisthe
responsibilityofeachstudent,whomustbeabletoconceive,carryoutandwriteup(asthethesis)a
significantbodyofworkinalogicalmanner.Allfacultyareavailablefordiscussionandconsultation.
Allstudentsareencouragedtoapplyforfellowshipsupportfromoutsideagencies,e.g.NIH,NSF,
HowardHughesMedicalInstitute,MarchofDimes,etc.
C.ThesisCommitteeMeetings
Itisrecommendedthatthestudentsmeetwiththeircommitteeseverysixmonths.Thecandidate
shouldprovidetheprogramwitha2030minutetalkshouldbegiveaswellasabriefwrittensummary
oftheprogressmadeonthestatedaimsgiventothecommitteeatleastoneweekpriortothemeeting.
Studentsarerequiredtomeetatleastonceeachyear.Thefollowingdocumentationisrequiredand
copiesmustbesubmittedtotheProgramAdministratorforinclusionintothestudentsfile:
dateofmeeting
studentswrittenreporttothethesiscommittee
thecommitteesresponses/recommendations
listofattendeesforeachmeeting
signaturesofstudentandcommitteechairman
D.Ph.D.Thesis
AfterpassingtheComprehensiveExamination,thestudententersPh.D.candidacy.Duringthe
followingyearsthestudentsperformresearchtowardsathesisdefense.Studentsmustgive
annualreportsontheprogressoftheirthesisresearchtotheCSDfacultyintheformof30
minuteseminars,andmeetatleastannuallywiththeirThesisCommittee.TheChairmanof
theThesisCommitteewillmeetwiththeGACtodiscussthestudentsprogressandwill
submitabriefwrittensummaryoftheoutcomeofeachmeetingwiththestudent.
Uponcompletion ofabodyoforiginal research that constitutes asignificant contribution ofnew
knowledgetothefieldofcellanddevelopmentalbiology,studentswillwriteaPh.D.thesiscontaining
thisinformation,anddefendthisdocumentatanoralexaminationscheduledbytheGraduateSchool.
Check with the Graduate School for current deadlines, thesis format requirements and required
paperworkpriortowritingthethesisandschedulingthedefense.
1.Guidelines
TherulesoftheUniversityofColoradoGraduateSchoolconcerningaPhDthesisareas
follows:Alldoctoralstudentsarerequiredtosubmitathesis(ordissertation)totheGraduateSchool
aspartialfulfillmentoftherequirementsofthedegreeofDoctorofPhilosophy.Theformandscopeof
thisthesisisdeterminedbythestudent,thethesisadvisor,theAdvisoryCommittee,andtheProgram.
Thethesisshouldbebaseduponoriginalinvestigationandshowingmaturescholarshipandcritical
judgmentaswellasfamiliaritywithtoolsandmethodsofresearch.Itmustbeessentiallyapprovedby
theexaminingcommitteebeforethefinalexaminationcanbetaken.
TheGraduatePrograminCellBiology,StemCellsandDevelopmentamplifiesthedefinitionofthe
thesisasfollows:
Thesuccessfulthesispresentsaproblemorientated,originalandsubstantive
investigation.Themethodologyandresultscontainedinthethesismustbeconclusive
13

andofquality.Thestandardsaretobethosemaintainedbyquality,peerreviewed
scientificjournals.Itistheexpectationoftheprogramthatthestudenthave1or
morefirstauthorpublicationssubmittedpriortothethesisdefense.
2.ThesisCommittee
Onceastudentisadmittedtocandidacy,he/sheshouldestablishaThesisCommitteewiththe
adviceofthethesisadvisorandtheDirectoroftheGraduateProgram.Thecommitteeneednotbethe
sameastheComprehensiveExamCommitteebutshouldbecomposedoffiveGraduateFaculty
members;atleastonemembermustbeoutsidetheprogramandthemajorityfromwithintheprogram.
Thethesisadvisorisavotingmemberofthiscommittee.Onefacultymemberoftheprogramshould
beselectedtoserveasachairoftheThesisCommittee.Thiscommitteecanbethesameasthe
ComprehensiveExamCommittee.
3.GraduateAdvisor
TheChairoftheThesisCommitteeservesastheadvisortothestudentandwillmonitorhis/her
progress.TheChairmustbeamemberoftheProgram.Itcannotbeemphasizedenough,however,that
eachstudentisresponsibleforhis/herownprogress.
4.GuidelinesforSupervisionofThesisWork
1.Sinceallstudentspresenttheirworkeachyear,allGraduateFacultyshouldfollowtheprogressof
allstudents.Whenconcernsarisetheyshouldbediscussedimmediatelywiththestudent,theThesis
Advisorand/orthestudentsThesisCommittee.
2.Studentsareencouragedtomeeteverysixmonths,butmustmeetatleastonceayear,withtheir
ThesisCommittee.StudentsmustsubmitawrittenupdateontheirprogresstotheCommitteeatleast
oneweekbeforetheCommitteemeeting.TheChairoftheCommitteeshouldprovidethestudentand
theProgramDirectorwithawrittensummaryofthestudentsprogressandrecommendationsofthe
thesisCommittee.Themeetingsshouldbedocumented(dateofmeeting,itemsdiscussed,committee
recommendations,listofattendees,signaturesofthestudentandcommitteechairman)andacopy
providedtotheProgramAdministratorforinclusionintothestudentsfile.TheThesisCommitteecan
recommendmorefrequentmeetingswhentheyfeelmorecarefulmonitoringiswarranted.
3.Whenthestudentandhis/herthesisadvisoragreetheworkforthethesishasbeencompleted,the
studentmustmeetwiththeThesisCommitteeandreceiveformalapprovaltobeginwritingthethesis.
5.PreparationofThesisandThesisDefense
1.TheStaffAssistantoftheGraduateSchool(currently,FranOsterberg)holdsseminarstwiceayear
ontheproperformattingofthethesis.
2.TheThesisCommitteemustformallyapprovethewrittenthesisbeforethefinalexaminationcanbe
taken.WrittenPhDthesisapprovalfromthechairoftheThesisCommitteeisrequiredpriorto
schedulingofthethesiswiththeGraduateSchool.TheThesisApprovalFormmaybeobtainedfrom
theprogramadministrator.Furthermore,thethesisadvisormustfindthethesisacceptablepriorto
submissiontotherestofthecommittee.Itisinexcusableforeveryoneconcernedifthestudentreaches
thepointofhis/herPhDthesisdefenseandencountersmajordifficultieswiththethesis.
14

3.Inadditiontocompletingthethesisdocument,priortothedefenseofthethesis,eachCSDstudent
mustsubmitaminimumofoneoriginalresearchmanuscriptforpublicationinordertoreceivethe
PhD.Thepapermustbefirstauthoredbythestudent,andrepresentacomponentofthestudents
overallthesiswork.Secondormiddleauthorshiporauthorshipofareviewarticleorchapterdoesnot
meetthisrequirement.
4.ArrangementsforthethesisdefensemustbemadeintheGraduateSchoolofficeatleastthreeweeks
inadvance.Theexaminationmustbetakenatleastthreeweekspriortothedateonwhichthedegreeis
tobeconferred.DegreesareconferredinMayandDecember.Thestudentmustberegisteredatthe
timeofthethesisdefense.Inaddition,acopyofthethesismustbegiventotheThesisCommitteeat
leasttwoweeksbeforethedefense.
4.Thethesisdefenseisthefinalexaminationofthethesisandrelatedtopics.Itincludesanoral
presentationofthesalientpointsoftheresearch,itsconclusionsanditsintegrationwiththerestofthe
field.TheoralpresentationwillbeconductedbytheThesisCommitteeandonlymembersofthe
GraduateFacultymaybepresent.Thefinaldecisionregardingtheresultofthethesisdefenseismade
bytheCommittee.
5.AllcorrectionstothewrittenthesisrequiredbytheThesisCommitteemustbecompletedwithin
thirtydaysfromthedateofthethesisdefense.Thesigned,writtendocumentmustbesubmittedtothe
GraduateSchoolatthattime.
6.Thestudentmustreceiveaffirmativevotesfromthemajorityofthecommittee.Theexamination
maybeattemptedonlyonce.Disqualificationofthethesisexaminationresultsindismissalfromthe
GraduateProgramwithoutadegree.
7.ThestudentisresponsibleforprovidingaboundcopyofthethesistotheThesisAdvisor,the
membersoftheThesisCommittee,theProgramandtheGraduateSchool.

15

IV.GraduateSchoolstandards
A.Credits.TheGraduateSchoolrequiresatleast30semesterhoursincoursework(rotationsand
ResearchCSBI7650countascoursehours)and30semesterhoursofthesisresearchforthePhD
(researchhourscannotbeaccumulateduntilthesemesterbeforetheComprehensiveExamispassed).
AllworkundertakenasagraduatestudentmustbeincompliancewiththeacademicCodeofHonor
(seeAppendixAofUCDAMCGraduateStudentHandbook).
B.Maintenanceofa3.0GPA.AllstudentsmustmaintainanaverageofBorbetterintheircourse
work.StudentsareexpectedtoearnaBorbetterinallrequiredcourses.Onlyinexceptional
circumstancesmayaBinarequiredcoursebeacceptable,asdeterminedbypetitiontotheGAC.
RequiredcoursescompletedwithagradeofbelowBcannotbecountedtowardsPhDrequirements.
C.Preliminaryexam.Inordertocontinueintheprogram,astudentmustpassPartsIandIIofthe
PreliminaryExamattheendofthefirstyear.IfthePreliminaryexamisfailed,thestudentsrecord
willbereviewedbytheGAC.Atthispointthestudentmaybeaskedtoretakepartoftheexam,the
entireexam,orleavethePhDtrainingprogram.
D.RemedialandDisciplinaryActions.StudentswhosecumulativeGPAfallsbelow3.0willbe
placedonAcademicProbationbytheGraduateSchool.ThestudentmustearnaGPAof3.0ineachof
his/hernexttwosemestersinordertoberemovedfromAcademicProbation.TheGraduateSchool
requiresthatafterastudentisputonacademicprobation,he/shemustmaintaina3.0inallsubsequent
semesters.FailingtomeeteitherconditionwillleadtoimmediatedismissalfromtheGraduateSchool.
ABorbelowinanyrequiredcourseisconsideredunsatisfactoryacademicprogressandmorethan
oneBorbelowisgroundsforimmediatedismissalfromtheProgram.
Agraduatestudentwhoreceivesanunsatisfactorygradeinacourse(aBorbelow)mayrepeat
thatcourseonceorsuccessfullycompleteanalternativeassignment,uponwrittenrecommendation
fromtheGACandapprovalbytheGraduateSchoolDean(providedthecoursehasnotbeen
previouslyappliedtowardadegree).Thetwogradesreceivedwillbeaveragedincalculatingthegrade
pointaverage,andallgradesreceivedwillappearonthestudentstranscript.Thecoursemaybe
countedonlyoncetowardsatisfyingtheunitrequirementforthedegree.
Aftertwosemesters,aGACmeetingwillbeheldtodeterminethestudentsprogress.Ifthe
studentscumulativeGPAis3.0orabove,thestudentwillberemovedfromprobation.Ifthestudents
cumulativeGPAisbelow3.0,thechairoftheThesisCommittee,thethesisadvisorandthestudent
willmeetwiththeCSDSteeringCommittee.TheSteeringCommitteewillmakeoneofthefollowing
determinations:
1.Thestudentisnotingoodacademicstandingandwillbeplacedonprobationagainfornot
morethan30days.
2.Thestudentisnotingoodacademicstandingandwillbereleasedfromtheprogram.
AllmeetingswillbethoroughlydocumentedandgiventotheProgramAdministratorforplacement
intothestudentsfile.

16

E.ChangeinThesisLab.Ifastudentleavesathesislabforanyreason,(butisstillconsideredbythe
CSDGACtobeingoodacademicstanding)thestudenthas1currentsemester(butnomore
than90days)torelocatetoanotherthesislabanddetermineanewthesisadvisorif
necessary.Itisthestudentsresponsibilitytolocateanotherthesislaband/oradvisor.
Withinthose90days,thestudentmustrotateforaminimumof6weeksinapotentialnew
advisorslab,sothatthefinaldecisiontojointhenewlabcanbemadewithinthe1
semester/90daywindow.
F.TimeLimitofPhDStudies.StudentshavesixyearsfromthetimetheyenterGraduateSchoolto
completeallrequirementsforthedegree.Continuationaftersixyearsrequirestheapproval
ofthestudentsThesisCommitteeandtheCSDSteeringCommittee.Itwillalsobe
necessarytopassasecondcomprehensiveexamination,similarincontenttothefirst,before
athesisdefensecanbescheduled.
G.Leaveofabsence.ItisthepolicyoftheCSDprogramtograntleaveofabsencesonlyunder
extremecircumstances.AformallettermustbesubmittedtotheGACandprogramdirector
explainingthereasonsaleaveofabsenceisnecessary.Beforetheleaveistakenitmustbeapproved
bytheGACandtheGraduateSchool.Afterastudentjoinsathesislab,thePImustalsoapprovethe
leaveofabsence.AspertherulesoftheGraduateSchool,only15daysofpaidleaveisguaranteed,
afterthattimeperioditisunderthediscretionoftheprogramandthePI.
V.ObligationsandRecordKeeping
A.Attendance.AllgraduatestudentsarerequiredtoattendtheweeklyCells,Developmentand
Cancer(CDC)seminars(usually,butnotalways,heldonWednesdaysatnoon)andspecialized
researchforums.Theseseminarsareamixtureoftalksbyinvitedspeakersandresearchreportsfrom
thefaculty,studentsandpostdoctoralfellowsinlaboratoriesoftheCellBiology,StemCellsand
DevelopmentProgramfaculty.
Thestudentsalsoorganizeabimonthlyjournalclub,andallCSDstudentsarerequiredtoparticipate.
Allgraduatestudentsarerequiredtoattendpostrotationalseminars,comprehensiveexaminations,
studentupdatepresentations,andThesisDefenseSeminarsgivenbyCSDprogramstudents.
Allnotebooks,originaldataandreagentsfromrotationalandthesisworkarethepropertyofthe
advisorandmustbeleftwiththeadvisoratthecompletionofthework.
Itisthestudentsresponsibilitytoregisterforcoursesinatimelymanneralllatefeesandfinance
chargeswillbetheresponsibilityofthestudent.
B.ColoradoResidency.FirstyearstudentswhoareUScitizensmustobtainaColoradoDrivers
LicenseatthetimeofarrivalattheUniversityofColoradoSchoolofMedicinetobeginthe
processofestablishingColoradoresidency.Ifresidencyhasnotbeenestablishedbythe
beginningofthesecondyear,thestudentisresponsibleforthenonresidentportionof
tuitionthatexceedstheresidentassessment.ThepaperworkforestablishingColorado
ResidencymustbefiledwiththeRegistrarpriortosecondyearregistration.
17

C.StudentsFiles.AfileforeachstudentwillbekeptbytheProgramAdministrator.Allrelevant
recordsshouldbegiventotheProgramAdministratorforthefiles,includingpublished
abstractsandpapers,notificationsofawardsandhonors,andcopiesofformsfiledwiththe
GraduateSchool.Thesefilesshouldreflectthetotalrecordofthestudentduringhis/her
entiregraduatecareer.Uponwrittenrequest,therecordsmaybeexaminedbythestudent.

VI.OtherCSDProgramevents
A.AnnualStudentResearchRetreat.Eachfall,thestudentshostanoutoftownretreatforthe
studentsandfacultyintheGraduatePrograminCellBiology,StemCellsandDevelopment.
Thepurposeoftheretreatistwofold:1)toprovideeveryonewiththeopportunitytoget
togetherandinteractonascientific/intellectuallevelsoastocultivatenewinteractionsand
strengthenexistingones;and2)toprovideanopportunityforincomingfirstyearCSDand
BiomedicalSciencesProgram(BSP)graduatestudents,and1stand2ndyearMSTPstudents
tobecomefamiliarwiththeresearchactivitiesandfacultywithintheCSDProgram.The
retreatisusuallyheldinSeptemberorOctober.Currentseniorstudents(2ndyearand
beyond)areexpectedtopresenttheirworkeitherviaaposteroratalk.
B.ParticipationinRecruitmentFunctions.DuringFebruary/Marcheachyear,prospectivestudent
applicantsvisitourprogramforinterviews.ItisintheProgramsbestinteresttoattractandretainthe
bestoftheseprospectivestudents.TodothisweneedthehelpofcurrentstudentsandCSDfaculty
whocanconvincetheseindividualsthatourProgramistheplacetobe!Whenasked,pleasebewilling
tospendsometimewithprospectivestudentsduringdinnersorotherfunctions.OurCSDProgram
canandhasflourishedwithyourirreplaceablehelp.
C.Descriptionofcommittees.Eachcommitteewithintheprogramhasastudentrepresentative.
Belowisadescriptionofthedutiesforeachposition:
Recruitment:Thiscommitteereviewssubmittedapplicationstothegraduateprogram,selects
candidatestointerviewinpersonorbytelephone,organizesrecruitmentweekend,andultimately
selectswhowillbeadmittedtotheprogram.Thestudentmembersparticipatefullyintheentire
process,andinparticularareinchargeofenlistingandorganizingthestudentbodytohelpwithboth
academicandsocialrecruitmentefforts.
Advising:TheGraduateAdvisoryCommitteehelpsstudentsmaintainprogresstowardtheirPh.D.
degree.Asamemberofthiscommittee,thestudentmemberhelpsdiscussstudentprogressandmaybe
recruitedtoand/oradvisethecommitteetotutorfirstyearstudentsinneed.Thiscommitteerequires
thatthestudentmemberbeadoctoralcandidate,i.e.,haspassedtheComprehensiveExam.Inaddition,
thestudentmemberoftheadvisingcommitteealsoservesontheSteeringCommitteeofthegraduate
program.
Curriculum:Thiscommitteediscussesthecurrentcurriculumandsuggestsandimplementschanges
inthebestinterestoftheprogramandstudents,includingbutnotlimitedtoselectionofAdvanced
Topicscoursestobeofferedeachacademicyearbasedonfacultyavailabilityandstudentinterests.

18

Membership:Thiscommitteeisresponsibleforfacultymembershipwithintheprogram.Thestudent
memberparticipatesintheestablishmentofguidelinesforfacultymembership,reviewscurrentfaculty
participation,andmakesrecommendationsconcerningnewfacultyapplicants.
Retreat:Everyfall,theprogramhasaretreat(typicallyovernight)inamountainlocation.Itistherole
ofthestudents(three)ontheRetreatCommitteetoorganizetheretreatwiththeoversightofaCSD
graduateprogramfacultyrepresentative.Withapredeterminedbudgetinmind,thestudentshavea
roleinselectingthelocationaswellasaninvitedspeaker.
SteeringCommittee:Thiscommitteeconsistsofthechairsofeachofthe6graduateprogram
committeesplusadditionalmembersasdeemedappropriatebythedirectoroftheprogram.
CommitteeofStudents:Consistsofthestudentmembersofthe5GraduateProgramCommittees
(Advising,Recruitment,Curriculum,MembershipandRetreat)andischairedbythestudentmember
oftheAdvisingCommittee.Thegoalofthiscommitteeistoorganizestudentactivitiesandprovidea
mechanismtodiscussstudentissuesthatcanbedirectedtotheSteeringCommitteeand/orprogram
director.

19

UCDAMC
GraduatePrograminCellBiology,StemCellsandDevelopment
I.

AdvanceTopicsinCellBiology,StemCellsandDevelopment,andelectivesofferedby
otherdepartmentsand/orprograms.

StudentsmusttakeatleastfourAdvancedTopics(CSDV7670,contentvariesyeartoyear)or
electives,beginninginyeartwo.
A.

AdvanceTopicsinCellBiologyStemCellsandDevelopment(CSDV7670)

EachCSDVcourseis12creditsandcomprises1530hrsofmeetingtimewithinasemester.In
general,oneCSDV7670coursewillbeofferedperyear.Studentsareencouragedtosubmitideasfor
CSDVAdvancedTopicsCoursesofspecialinteresttothemtotheCurriculumCommittee.Students
whoareintheprogramforlongerthanfiveyearsarerequiredtotakeoneadditionalAdvancedTopic
peryearuntilgraduation.Thisrequirementhasbeenineffectsincethe199192academicyear.
B.

Electivesofferedbyotherdepartments/programs.

Afewelectivesarehighlightedhere,butcourseschangeyearly.Thebestresourceforcourseofferings
willbefoundontheregistrarswebsitewhenyouregisterforeachupcomingsemester.However,
belowwelistseveralfrequentlyofferedelectivesforadvancedgraduatestudents.
IDPT7646

TissueBiologyandDiseaseMechanism

J.Hooper

IMMU7630

MOLB7800

NRSC7615

OverviewofImmunology

J.Cohen

AdvancedTopicsinMolecularBiology

J.Kieft

DevelopmentalNeurobiology

A.Ribera/K.Artinger

PHCL7606

ReceptorsandCellSignaling

M.DellAcqua

CANB 7600

C.

CancerBiology

S.Nordeen

IndependentStudiesinCellandDevelopmentalBiology(CSDV7850)

IndependentStudyistoaccommodatestudentswhowishto(1)takeaProfessionalSchool
Courseforcreditand(2)gainadefinedexpertisewithafacultymentorotherthantheirthesisadvisor.
ConsentofthefacultymemberofferingtheIndependentStudyandtheProgramDirectorarerequired.

20

CellBiology,StemCellsandDevelopmentGraduateProgram
CurrentstudentsasofSeptember2011
STUDENT

BrittanyAllen
FrancieBarron
YingZhang
AaronHuebner
JohnSchiel
ZeljkoDvanajscak
DongyingLi
AlexBlasky
ShiriGutman
BradKubick
DavalynPowell
JenniferIkle
JinxiangDai
JasonDinella
JonathanWilde
HeatherRay
DavidCastillo
BrianBayless
CourtneyBetts
MaryPinter
DianeGumina
AndrewWeems

STARTYEAR

THESIS
ADVISOR

PRELIMS

COMPS

THESIS
COMMITTEE
CHAIR

2005
2006
2006
2006(BSP)
2007
2007(MSTP)
2008

Reyland
Clouthier
Niswander
Roop
Jones
Appel
Prekeris

6/06
6/07
6/07
6/07
6/08
6/08
6/08

9/07
9/08
11/08
9/08
12/09
7/10
9/10

Schedin
Niswander
Hooper
Niswander
Doctor
Finger
Doctor

2008
2008(BSP)
2008(BSP)
2008(BSP)
2008(MSTP)
2009
2010
2010
2010(BSP)
2010(BSP)
2010(BSP)
2011
2011
2011
2011

Appel
Evans
Roop
Artinger
Clouthier
Macklin
Koch
Niswander
Niswander
Barlow
Pearson

6/08
6/08
6/08
6/08
6/07
6/10
6/11
6/11
6/11
6/11
6/11

12/10
9/10
11/10
9/10
9/09

Reyland
Wang
Clouthier
Appel

21

CSD FACULTY ROSTER AUGUST 2011

Faculty

Primary Dept

CSD Students

Bruce Appel
Kristin Artinger
Vivek Balasubramanian
Linda Barlow
Lynne Bemis
Brad Bendiak
Steve Britt
John Caldwell
David Clouthier
Richard Davis
Stijn DeLanghe
Brian Doctor
Tobias Eckle
Holger Eltzschig
Tom Evans
Tom Finger
Jim Hagman
Joan Hooper
Kathryn Howell
John Hutton
Karen King
Peter Koch
Maranke Koster
Shilong Lu
Wendy Macklin
Jim McManaman
Lee Niswander
Chad Pearson
Karl Pfenninger
Rytis Prekeris
Yosef Refaeli
Diego Restrepo
Jane Reusch
Mary Reyland
Sue Reynolds
Dennis Roop
Pepper Schedin
Geeta Sharma
Trevor Williams
Virginia Winn
Xiaojing Wang

Pediatrics
Craniofacial Biology
Pulmonary Medicine, Pediatrics
Cell & Developmental Biology
Medical Oncology/Medicine
Cell & Developmental Biology
Cell & Developmental Biology
Cell & Developmental Biology
Craniofacial Biology
Biochemistry
Immunology/Peds
Gastroentoerology/Medicine
Anesthesiology
Anesthesiology
Cell & Developmental Biology
Cell & Developmental Biology
Pediatrics
Cell & Developmental Biology
Cell & Developmental Biology
Barbara Davis Center/Pediatrics
Orthopedics
Dermatology
Dermatology
Otolaryngology
Cell & Developmental Biology
Obstetrics and Gynecology/Medicine
Pediatrics
Cell & Developmental Biology
Pediatrics
Cell & Developmental Biology
Dermatology
Cell & Developmental Biology
Endocrinology/Medicine
Craniofacial Biology
Immunology/Peds
Dermatology
Medical Oncology/Medicine
Physiology
Craniofacial Biology
Obstetrics and Gynecology/Medicine
Pathology

Blasky, Dvanajscak
Powell

41 active faculty members. 37 training faculty

22

Castillo

Hyndman, Ikle

Gutman

Dinella
Dai
Zhang, Wilde, Ray
Bayless
Schiel, Li

Adwan, Allen
Huebner, Kubick

UCDAMC
GraduatePrograminCellBiology,StemCellsandDevelopment
20112012CommitteeMembershipRoster
LindaBarlow,Director
BruceAppel,AssociateDirector
RecruitmentCommittee
KristinArtinger,Chair
ChadPearson
TBD
BSPAdmissionsCommittee
MarankeKoster
GraduateAdvisoryCommittee
TomEvans,Chair
JoanHooper
BrianDoctor
ZeljkoDvanajscak
CurriculumCommittee
LeeNiswander,Chair
RytisPrekeris
MaryReyland
DavalynPowellandDongyingLi
MembershipCommittee
JimMcManaman,Chair
JoanHooper
DavidClouthier
KarenKing
VivekBalasubrananian
JohnSchiel
CSD2011RetreatCommittee
LindaBarlow,Chair
JennIkleandDavalynPowell
CDCSeminarSeriesRepresentatives
DavidClouthierCFB
BruceAppelPeds
SteveBrittCDB
23

STEERINGCOMMITTEE
LindaBarlow
BruceAppel
SteveBritt
LeeNiswander
JimMcManaman
MarankeKoster
KristinArtinger
WendyMacklin
DennisRoop
CSD2012RetreatCommittee
TBD,Chair

24

UCDAMC
GraduatePrograminCellBiology,StemCellsandDevelopment
DegreesconferredasofSeptember2007
Student

Dateofthesis
defense

ThesisAdvisor

Currentposition

AllisonVigers(PhD)

2/5/92

Selitrennikoff

SeniorResearcher
CUBoulder
DeptofMolecular,Cellularand
DevelopmentalBiology

BrentWright(MD,PhD)

5/12/92

Scholey

Resident,UCDAMC

CorinnaBurger(PhD)

8/25/94

Ribera

AssistantProfessor
UniversityofFlorida,
DeptofNeurology

KathrynLohse(PhD)

8/2/94

Pfenninger

JeffCrosby(PhD)

10/21/94

Howell

DirectorofInflamationResearch,
ISISPharmaceuticals,SanDiego

HilarySrere(PhD)

1/17/95

Martin

MarketingManager,
BioRadLaboratories

PaulAwald(PhD)

5/19/95

Selitrennikoff

Postdoc,MIT

ColinMonks(PhD)

6/19/95

Kupfer

President
IntelligentImagingInnovations
Denver,CO

KurtGriffin(MD,PhD)

3/96

Frerman

IntramuralResearcher
NIH/NICHD
SectiononEndocrinology&
Genetics

BarbaraBiery(PhD)

3/19/96

Goodman

Technicalwriter,
LifeTechnologies

LeslieStoneRoy(PhD)

1/24/96

Finger

SeniorResearcher
ColoradoStateUniversity
Dept.ofBiomedicalSciences

StevenJones(PhD)

5/27/97

Howell

AssociateDirectorofDiscovery
&AnimalBiology
ElusysTherapeuticsInc.,
NewJersey

25

ToniaVonOhlen(PhD)

10/8/97

Hooper

NicholeD.Schueck(MS)

11/14/97

Koeller

Maria(Johnson)Curtis(PhD)

4/9/98

Granholm

DeniseSalazarRogers(PhD)

4/29/98

Frerman

AssocScientificDirector,
QuestDiagnostics

HilaryChouinard(MS)

12/98

Ribera

PlannedParenthood,Denver

KennethKosten(MS)

12/98

Garcea

RandyTaylor(MD,PhD)

7/12/99

Howell

ENTPhysicianinPrivatePractice

MarkNordeen(PhD)

10/19/99

Caldwell

AssociateDeanforArts&
Sciences
CentralWyomingCollege,
Riverton,Wyoming

EricBush(PhD)

11/03/99

Perryman

ChristineWu(PhD)

3/12/01

Howell/Neville

AssistantProfessor
UCDAMC
DeptofPharmacology

MizunoSugawara(MS)

3/19/01

Sussel

Homemaker,Japan

FangtianHuang(MS)

6/1/01

Sorkin

RussBowler(MD,PhD)

6/21/01

Crapo

AssociateProfessor,
UniversityofColoradoHealth
SciencesCenter,
DeptofMedicine

KeithMikule(PhD)

5/17/01

Pfenninger

ArQuleBiomedicalInstitute,Inc.,
Norwood,Massachusetts

JoshHall(MD,PhD)

7/27/01

Finger

Resident
UnivofCalifornia,SanDiego
DeptofPsychiatry

LuisMiranda(PhD)

5/2/02

Franzusoff

JamesWitowsky(PhD)

11/19/02

Johnson

26

AssistantProfessor
KansasStateUniversity,
DivisionofBiology

ResearchInstituteofMolecular

PathologyVienna,Austria
CindyYee(PhD)

4/23/03

Finger

ScottBarbee(PhD)

5/5/04

Evans

JenniferGillette(PhD)

5/15/04

NielsenPreiss

AlexLublin(PhD)

11/18/04

Evans

Postdoc,Mt.SinaiHospital

JayGatlin(PhD)

12/15/04

Pfenninger

Postdoctoralresearcher
Univ.ofNorthCarolina,Chapel
Hill
Dept.ofBiology

DavidKent(PhD)

7/19/04

Hooper

Postdoc,UnivUtah

DeniseBirkholz(PhD)

7/15/05

Britt

Postdoctoralfellow
UCDAMC
Dept.ofCraniofacialBiology

MikeHumphries(PhD)

9/21/05

Reyland

ChristinaChao(PhD;MSTP)

3/6/06

Sussel

Postdoctoralresearcher,
ArrayBioPharma,Boulder
UnivofColoradoSchoolof
Medicine

JamesEarl(Ph.D;MSTP)

1/13/06

Britt

NickosSophos(MS)
MalindaODonnell(MS)
AgneTaraseviciute(Ph.D;MSTP)

7/06
2007
2008

Reyland
Garcea
P.Jones

RoslynBauer(PhD)

2008

J.Hutton

GlennSimon(PhD)

2008

R.Prekeris

DanielleHarlow(PhD)

2009

L.Barlow

JianJing(PhD)

2009

R.Prekeris

IreneChoi(MS)
DavidMcKean(PhD)

2009
2010

D.Roop
L.Niswander

LaiKuanGoh(PhD)

2010

A.Sorkin

27

ResearchScientist,
Genentech
SanFrancisco,CA
AssistantProfessor,Universityof
Denver,DeptofBiological
Sciences
Postdoctoralfellow
NIH/NICHD
SectiononOrganelleBiology

UnivofColoradoSchoolof
Medicine

Univ.ofColoradoSchoolof
Medicine
JoVE(JournalofVisualized
Experiements)
Postdoctoralfellow,Univof
ColoradoSchoolofMedicine
Postdoctoralfellow,Univof
ColoradoSchoolofMedicine
Postdoctoralfellow,National
JewishMedResearchCenter
Postdoctoralfellow,Harvard
MedicalSchool

HaNguyen(PhD)
TariqAdwan(PhD)

2011
2011

L.Barlow
M.Reyland

28

HanoiMedicalUniversity