Académique Documents
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1 Etymology
2 EVOLUTIONARY HISTORY
carnassials are attened
all living bears. Species of Ursavus subsequently entered North America, together with Amphicynodon and
Additionally, members of this family possess posteriorly Cephalogale, during the early Miocene (2118 Mya).
oriented M2 postprotocrista molars, elongated m2 mo- Members of the living lineages of bears diverged from
lars, and a reduction of the premolars.
Ursavus between 15 and 20 Mya ago,[10][11] likely via the
Modern bears comprise eight species in three subfam- species Ursavus elmensis. Based on genetic and morphoilies: Ailuropodinae (monotypic with the giant panda), logical data, the Ailuropodinae (pandas) were the rst to
Tremarctinae (monotypic with the spectacled bear), and diverge from other living bears about 19 Mya ago, alof this group have been found before
Ursinae (containing six species divided into one to three though no fossils
[12]
about
5
Mya.
genera, depending on the authority).
2.1
Fossil bears
The New World short-faced bears (Tremarctinae) differentiated from Ursinae following a dispersal event
into North America during the mid Miocene (about 13
Mya).[12] They invaded South America (~1 Ma) following formation of the Isthmus of Panama.[13] Their earliest fossil representative is Plionarctos in North America (~ 102 Ma). This genus is probably the direct ancestor to the North American short-faced bears (genus
Arctodus), the South American short-faced bears (Arctotherium), and the spectacled bears, Tremarctos, represented by both an extinct North American species (T.
oridanus), and the lone surviving representative of the
Tremarctinae, the South American spectacled bear (T.
ornatus).
The subfamily Ursinae experienced a dramatic proliferation of taxa about 5.34.5 Mya ago, coincident with major environmental changes; with the rst members of the
genus Ursus also appearing around this time.[12] The sloth
bear is a modern survivor of one of the earliest lineages
to diverge during this radiation event (5.3 Mya); it took
on its peculiar morphology, related to its diet of termites
and ants, no later than by the early Pleistocene. By 34
Mya ago, the species Ursus minimus appears in the fossil
record of Europe; apart from its size, it was nearly identical to todays Asiatic black bear. It is likely ancestral
to all bears within Ursinae, perhaps aside from the sloth
bear. Two lineages evolved from U. minimus: the black
bears (including the sun bear, the Asiatic black bear, and
the American black bear); and the brown bears (which includes the polar bear). Modern brown bears evolved from
3
U. minimus via Ursus etruscus, which itself is ancestral to
both the extinct Pleistocene cave bear and todays brown
and polar bears. Species of Ursinae have migrated repeatedly into North America from Eurasia as early as 4
Mya during the early Pliocene.[14]
3 Classication
The fossil record of bears is exceptionally good. Direct ancestor-descendent relationships between individual species are often fairly well established, with sucient
intermediate forms known to make the precise cut-o between an ancestral and its daughter species subjective.[15]
Other extinct bear genera include Agriarctos, Indarctos, and Agriotherium (sometimes placed within
hemicyonids).
2.2
Family Ursidae
CLASSIFICATION
Ailuropoda microta
Ailuropoda wulingshanensis
Subfamily Ailuropodinae
Ailurarctos
Ailurarctos lufengensis
Ailurarctos yuanmouenensis
Ailuropoda (pandas)
Ailuropoda baconi
Ailuropoda fovealis
Ailuropoda melanoleuca - giant
panda
Ailuropoda
melanoleuca
melanoleuca; giant panda
Ailuropoda melanoleuca qinlingensis; Qinling panda
Subfamily Tremarctinae
Plionarctos
Plionarctos edensis
Plionarctos harroldorum
Tremarctos (spectacled bears)
Tremarctos ornatus - spectacled
bear
Tremarctos oridanus - Florida
spectacled bear
Arctodus
Arctodus simus - giant short-faced
bear
Arctodus pristinus
Arctotherium
Arctotherium angustidens
Arctotherium bonariense
Arctotherium brasilense
Arctotherium latidens
Arctotherium tarijense
Arctotherium vetustum
Arctotherium wingei
Subfamily Ursinae
Ursavus
Ursavus brevirhinus
Ursavus depereti
Ursavus elmensis
Ursavus pawniensis
Ursavus primaevus
Ursavus tedfordi
Indarctos
Indarctos anthraciti
Indarctos arctoides
Indarctos atticus
Indarctos nevadensis
Indarctos oregonensis
Indarctos salmontanus
Indarctos vireti
Indarctos zdanskyi
Agriotherium
Agriotherium inexpetans
Agriotherium schneideri
Agriotherium sivalensis
Melursus
Melursus ursinus - sloth bear
Melursus ursinus inornatus; Sri
Lankan sloth bear
Melursus ursinus ursinus; Indian
sloth bear
Helarctos
Helarctos malayanus - sun bear
Helarctos malayanus malayanus
Helarctos malayanus euryspilus;
3
Borneo sun bear
Ursus
Ursus rossicus
Ursus sackdillingensis
Ursus minimus
Ursus thibetanus - Asian black bear
Ursus thibetanus formosanus; Formosan black bear
Ursus thibetanus gedrosianus
Ursus
thibetanus
japonicus;
Japanese black bear
Ursus thibetanus laniger
Ursus thibetanus mupinensis
Ursus thibetanus thibetanus
Ursus
thibetanus
ussuricus;
Manchurian black bear or Ussuri black bear
Ursus abstrusus
Ursus americanus - American
black bear
Ursus americanus altifrontalis,
Olympic black bear
Ursus americanus amblyceps, New
Mexico black bear
Ursus americanus americanus,
Eastern black bear
Ursus americanus californiensis,
California black bear
Ursus americanus carlottae, Haida
Gwaii black bear or Queen Charlotte black bear
Ursus americanus cinnamomum,
cinnamon bear
Ursus americanus emmonsii, glacier
bear
Ursus americanus eremicus, Mexican black bear
Ursus americanus oridanus,
Florida black bear
Ursus americanus hamiltoni, Newfoundland black bear
Ursus americanus kermodei, Kermode bear or spirit bear
Ursus
americanus
luteolus,
Louisiana black bear
Ursus americanus machetes, West
Mexico black bear
Ursus americanus perniger, Kenai
black bear
Ursus americanus pugnax, Dall
black bear
Ursus americanus vancouveri, Vancouver Island black bear
Ursus etruscus
CLASSIFICATION
4.1
Morphology
Biology
4.1
Morphology
4.1.1 Dentition
Despite being quadrupeds, bears can stand and sit similarly to
humans.
Bears are generally bulky and robust animals with relatively short legs. They are sexually dimorphic with regard to size, with the males being larger.[23][24] Larger
species tend to show increased levels of sexual dimorphism in comparison to smaller species,[24] and where a
species varies in size across its distribution, individuals
from larger-sized areas tend also to vary more. Bears are
the most massive terrestrial members of the order Carnivora. Some exceptional polar bears and Kodiak bears
(a brown bear subspecies) have been weighed at over 750
4.2
4 BIOLOGY
4.3
Behaviour
4.7
Winter dormancy
dependent upon the food supply available to the growing individual. The females of smaller species may have
young in as little as two years, whereas the larger species
may not rear young until they are four or even nine years
old. First breeding may be even later in males, where
competition for mates may leave younger males without
access to females.[26]
The bears courtship period is very brief. Bears in northern climates reproduce seasonally, usually after a period of inactivity similar to hibernation, although tropical species breed all year round. Cubs are born toothless, blind, and bald. The cubs of brown bears, usually
born in litters of one to three, will typically stay with the
mother for two full seasons. They feed on their mothers
milk through the duration of their relationship with their
mother, although as the cubs continue to grow, nursing
becomes less frequent and cubs learn to begin hunting
with the mother. They will remain with the mother for
about three years, until she enters the next cycle of estrus
and drives the cubs o. Bears will reach sexual maturity
in ve to seven years. Male bears, especially polar and
brown bears, will kill and sometimes devour cubs born to
another father to induce a female to breed again. Female
bears are often successful in driving o males in protection of their cubs, despite being rather smaller.
4.6
Breeding
in the winter, a belief supported by a number of scientic studies.[45][46] While many bear species do go into a
physiological state often colloquially called hibernation
or "winter sleep", it is not true hibernation. In true hibernators, body temperatures drop to near ambient and
American black bears mating
heart rates slow drastically, but the animals periodically
rouse themselves to urinate or defecate and to eat from
variable, both between and within species. Sexual ma- stored food. The body temperature of bears, on the other
turity is dependent on body condition, which is in turn hand, drops only a few degrees from normal, and the heart
10
Wojtek the bear with a Polish soldier: During the Battle of Monte
Cassino, Wojtek helped move ammunition.
Bear danger area closure sign of a type used in Denali National
Park, Alaska
bear, sloth bear, and brown bear, are dangerous to humans, especially in areas where they have become used to
people. All bears are physically powerful and are likely
capable of fatally attacking a person, but they, for the
most part, are shy, are easily frightened and will avoid
humans. Injuries caused by bears are rare, but are often widely reported.[48] The danger that bears pose is often vastly exaggerated, in part by the human imagination.
However, when a mother feels that her cubs are threatened, she will behave ferociously. It is recommended to
give all bears a wide berth because they are behaviorally
unpredictable.
Where bears raid crops or attack livestock, they may
come into conict with humans.[49][50] These problems
may be the work of only a few bears, but they create a
climate of conict, as farmers and ranchers may perceive
all losses as due to bears and advocate the preventive removal of all bears.[50] Mitigation methods may be used to
reduce bear damage to crops, and reduce local antipathy
towards bears.[49]
Laws have been passed in many areas of the world to protect bears from habitat destruction. Public perception of
bears is often very positive, as people identify with bears
due to their omnivorous diets, ability to stand on two legs,
and symbolic importance,[51] and support for bear protection is widespread, at least in more auent societies.[52]
In more rural and poorer regions, attitudes may be more
6.2
Culture
6.1
Names
11
fairy tale "Morozko", whose arrogant protagonist Ivan
tries to kill a mother bear and her cubsand is punished
and humbled by having his own head turned magically
into a bears head and being subsequently shunned by human society.
"The Brown Bear of Norway" is a Scottish fairy tale
telling the adventures of a girl who married a prince magically turned into a bear, and who managed to get him
back into a human form by the force of her love and after
many trials and diculties. In the 1970s, this story was
adapted into the East German fantasy lm The Singing
Ringing Tree and broadcast on British television.
Evidence of bear worship has been found in early Chinese
and Ainu cultures, as well (see Iomante). Korean people
in their mythology identify the bear as their ancestor and
symbolic animal. According to the Korean legend, a god
imposed a dicult test on a she-bear; when she passed it,
the god turned her into a woman and married her.
In Russian and other Slavic languages, the word for bear, Legends of saints taming bears are common in the Alpine
medved (), and variants or derivatives, such as zone. In the arms of the bishopric of Freising, the bear
is the dangerous totem animal tamed by St. Corbinian
Medvedev, are common surnames.
and made to carry his civilised baggage over the mounThe Irish family name McMahon means Son of Bear
tains. A bear also features prominently in the legend of
in Irish.
St. Romedius, who is also said to have tamed one of these
In East European Jewish communities, the name Ber animals and had the same bear carry him from his her()Yiddish cognate of Bearhas been attested as mitage in the mountains to the city of Trento.
a common male rst name, at least since the 18th century,
Similar stories are told of Saint Gall and Saint Columand was, among others, the name of several prominent
banus.
rabbis. The Yiddish Ber is still in use among Orthodox
Jewish communities in Israel, the US, and other coun- This recurrent motif was used by the Church as a symbol
tries. With the transition from Yiddish to Hebrew un- of the victory of Christianity over paganism.[59] In the
der the inuence of zionism, the Hebrew word for bear, Norse settlements of northern England during the 10th
dov (), was taken up in contemporary Israel and is at century, a type of hogback grave cover of a long narrow
present among the commonly used male rst names in block of stone, with a shaped apex like the roof beam of a
long house, is carved with a muzzled, thus Christianised,
that country.
bear clasping each gable end. Though the best collec"Ten Bears" (Paruasemana) was the name of a welltion of these is in the church at Brompton, North Yorkknown 19th century chieftain among the Comanche.
shire,[60] their distribution ranges across northern EngAlso among other Native American tribes, bear-related
land and southern Scotland, with a scattered few in the
names are attested.
north Midlands and single survivals in Wales, Cornwall,
and Ireland; a late group is found in the Orkney Islands.
Bears are a popular feature of many childrens stories, including Goldilocks and "The Story of the Three Bears",
For bears in mythology, see Jean de l'Ours, Arcturus, the Berenstain Bears, and Winnie the Pooh.
Ursa Major, and Berserker.
En uheldig bjrnejakt (An Unfortunate Bear Hunt)
by Theodor Kittelsen.
There is evidence of prehistoric bear worship. Anthropologists such as Joseph Campbell have regarded this as
Onikuma from Ehon Hyaku Monogatari
a common feature in most of the shing and hunting According to his hagiography, a bear killed Saint
tribes. The prehistoric Finns, along with most Siberian
Corbinian's pack horse on the way to Rome, so the
peoples, considered the bear as the spirit of ones foresaint commanded it to carry his load. Once he arfathers. This is why the bear (karhu) was a greatly rerived in Rome, however, he let the bear go.
spected animal, with several euphemistic names (such as
otso, mesikmmen and kontio). The bear is the national
The saddled bear of St. Corbinian" the emblem
animal of Finland.
of Freising, here incorporated in the arms of Pope
6.2
Benedict XVI
12
6 CULTURE
6.3
Symbolic use
The British Lion, the Persian Cat and the Russian Bear (see The
Great Game)
Also, bear, "bruin", or specic types of bears are popular nicknames or mascots, for example, for sports teams
(Bayern Munich, Chicago Bears, California Golden
Bears, UCLA Bruins, Boston Bruins); and a bear cub
called Misha was mascot of the 1980 Summer Olympics
in Moscow, Soviet Union.
The ag of California
The physical attributes and behaviours of bears are commonly used in gures of speech in English.
In the UK, the bear and sta has long featured on the
heraldic arms of Warwickshire county.[61]
In gay slang, the term "bear" refers to male individuals who possess physical attributes much like a bear,
13
such as a heavy build, abundant body hair, and com- Conservation.[65] Speciality organizations for each of the
monly facial hair.
eight species of bears worldwide include:
A bear hug is typically a tight hug that involves wrapping ones arms around another person, often leaving that persons arms immobile.
Bear tracking in the old Western states of the U.S.
and, to this day, in the former Dakota Territory, the
expression you ain't just a bear trackin'" is used to
mean you ain't lying or thats for sure. This expression evolved as an outgrowth of the experience
pioneer hunters and mountainmen had when tracking bear. Bears often lay down false tracks and are
notorious for doubling back on anything tracking
them. If you are not following bear tracks, you are
not following false trails or leads in your thoughts,
words or deeds.
In Korean culture, a person is referred to as being
like a bear when they are stubborn or not sensitive to what is happening around their surroundings.
Used as a phrase to call a person stubborn bear.
The Bible compares King David's bitter warriors,
who ght with such fury that they could overcome
many times their number of opponents, with a
bear robbed of her whelps in the eld (2 Samuel
17:8 s:Bible (King James)/2 Samuel#Chapter 17).
The phrase a bereaved bear () , derived
from this Biblical source, is still used in the literary
Hebrew of contemporary Israel.
6.5
Teddy bears
8 See also
List of fatal bear attacks in North America
List of ctional bears
List of individual bears
Ursa minor
Ursari
9 References
[1] Slovakia warns of tipsy bears. Archived from the original on 2010-11-18. Retrieved 2008-11-11.
Two authoritative organizations for seeking scientic information on bear species of the world are the International Association for Bear Research & Management,
also known as the International Bear Association (IBA);
and the Bear Specialist Group of the Species Survival
Commission, a part of the International Union for the
Conservation of Nature. These organizations focus on
the species natural history, management, and conservation.
Other organizations exist to further wild bear education
and conservation. Bear Trust International works for
wild bears and other wildlife through four core program
initiatives: 1) Conservation Education, 2) Wild Bear
Research, 3) Wild Bear Management, and, 4) Habitat
[4] hildebrand.raa.se
[5] Votruba, Martin. Bears. Slovak Studies Program. University of Pittsburgh. Archived from the original on 201011-18. Retrieved 2009-03-12.
[6] Welsey-Hunt, G.D. & Flynn, J.J. (2005). Phylogeny of
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[7] Wang, Xiaoming, Malcolm C. McKenna, and Demberelyin Dashzeveg (2005). Amphicticeps and Amphicynodon (Arctoidea, Carnivora) from Hsanda Gol Formation,
Central Mongolia and Phylogeny of Basal Arctoids with
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14
[8] Kemp, T.S. (2005). The Origin and Evolution of Mammals. Oxford University Press. ISBN 978-0-19-8507604.
[9] Wang Banyue and Qiu Zhanxiang (2005). Notes
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116124.
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Archived from
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[10] Waits, Lisette (1999).
Rapid radiation events in
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[11] Pges, Marie (2008). Combined analysis of fourteen nuclear genes renes the Ursidae phylogeny
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[12] Krause, J.; Unger, T.; Noon, A.; Malaspinas, A.;
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[13] Soibelzon, L. H.; Tonni, E. P.; Bond, M. (October 2005). The fossil record of South American
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[14] Qiu Zhanxiang (2003). Dispersals of Neogene Carnivorans between Asia and North America (PDF). Bulletin
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[15] Kurtn, B., 1995. The cave bear story: life and death of
a vanished animal, Columbia University Press, ISBN 0231-10361-1
[16] Lindburg, Donald G. (2004). Giant Pandas: Biology and
Conservation, pp. 79. University of California Press,
ISBN 0-520-23867-2
[17] Olaf R. P. Bininda-Emonds. Phylogenetic Position of
the Giant Panda. In Lindburg, Donald G. (2004) Giant
Pandas: Biology and Conservation, pp. 1135. University
of California Press, ISBN 0-520-23867-2
[18] Kutschera, Verena (2014). Bears in a forest of gene trees:
Phylogenetic inference is complicated by incomplete lineage sorting and gene ow. Molecular Biology and Evolution 31: 20042017. doi:10.1093/molbev/msu186. Retrieved 17 November 2014.
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15
[55] Bear meat poisoning in Siberia. BBC News. 1997-1221. Retrieved 2006-10-04.
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[56] Finnish Food Safety Authority: Bear meat must be inspected before serving in restaurants. Retrieved 200610-04.
[57] http://www.constellationsofwords.com/Constellations/
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16
11
10
Further reading
11
External links
EXTERNAL LINKS
17
12
12.1
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