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For other uses, see Potato (disambiguation). nally referred to a type of sweet potato although the two
Spud redirects here. For other uses, see Spud (disam- plants are not closely related; in many of the chronicles
biguation). detailing agriculture and plants, no distinction is made
between the two.[14] The 16th-century English herbal-
ist John Gerard used the terms bastard potatoes and
The potato is a starchy, tuberous crop from the perennial
nightshade Solanum tuberosum. The word potato may Virginia potatoes for this species, and referred to sweet
[15]
[2] potatoes as common potatoes. Potatoes are occa-
refer either to the plant itself or to the edible tuber.
In the Andes, where the species is indigenous, there sionally referred to as Irish potatoes or white potatoes
are some other closely related cultivated potato species. in the United States, to distinguish them from sweet pota-
[15]
Potatoes were introduced outside the Andes region ap- toes.
proximately four centuries ago,[3] and have since become The name spud for a small potato comes from the dig-
an integral part of much of the worlds food supply. It ging of soil (or a hole) prior to the planting of potatoes.
is the worlds fourth-largest food crop, following maize, The word has an unknown origin and was originally (c.
wheat, and rice.[4] The green leaves and green skins of 1440) used as a term for a short knife or dagger, proba-
tubers exposed to the light are toxic. bly related to Dutch spyd or the Latin spad-" a word root
Wild potato species can be found throughout the Amer- meaning sword"; cf. Spanish espada, English spade
icas from the United States to southern Chile.[5] The and spadroon. The word spud traces back to the 16th
potato was originally believed to have been domesti- century. It subsequently transferred over to a variety of
cated independently in multiple locations,[6] but later ge- digging tools. Around 1845, the name transferred to the
netic testing of the wide variety of cultivars and wild tuber itself.[16] The origin of the word spud has erro-
species proved a single origin for potatoes in the area neously been attributed to a 19th-century activist group
dedicated to keeping the potato out of Britain, calling it-
of present-day southern Peru and extreme northwest-
ern Bolivia (from a species in the Solanum brevicaule self The Society for the Prevention of an Unwholesome
Diet (S.P.U.D.).[16] It was Mario Pei's 1949 The Story of
complex), where they were domesticated approximately
7,00010,000 years ago.[7][8][9] Following centuries of Language that can be blamed for the words false origin.
Pei writes, the potato, for its part, was in disrepute some
selective breeding, there are now over a thousand dif-
ferent types of potatoes.[8] Over 99% of the presently centuries ago. Some Englishmen who did not fancy pota-
toes formed a Society for the Prevention of Unwholesome
cultivated potatoes worldwide descended from varieties
that originated in the lowlands of south-central Chile, Diet. The initials of the main words in this title gave rise
which have displaced formerly popular varieties from the to spud. Like most other pre-20th century acronymic
Andes.[10][11] origins, this is false.[16]
1 Etymology
1
2 3 GENETICS
3 Genetics
There are about 5,000 potato varieties worldwide. Three
thousand of them are found in the Andes alone, mainly
in Peru, Bolivia, Ecuador, Chile, and Colombia. They
belong to eight or nine species, depending on the tax-
onomic school. Apart from the 5,000 cultivated vari-
eties, there are about 200 wild species and subspecies,
many of which can be cross-bred with cultivated vari-
eties. Cross-breeding has been done repeatedly to trans-
Russet potatoes
fer resistances to certain pests and diseases from the gene
pool of wild species to the gene pool of cultivated potato
species. Genetically modied varieties have met pub-
Potato plants are herbaceous perennials that grow about lic resistance in the United States and in the European
[19][20]
60 cm (24 in) high, depending on variety, with the leaves Union.
dying back after owering, fruiting and tuber forma- The major species grown worldwide is Solanum tubero-
tion. They bear white, pink, red, blue, or purple owers sum (a tetraploid with 48 chromosomes), and modern
with yellow stamens. In general, the tubers of varieties varieties of this species are the most widely cultivated.
with white owers have white skins, while those of vari- There are also four diploid species (with 24 chromo-
eties with colored owers tend to have pinkish skins.[17] somes): S. stenotomum, S. phureja, S. goniocalyx, and
Potatoes are mostly cross-pollinated by insects such as S. ajanhuiri. There are two triploid species (with 36
bumblebees, which carry pollen from other potato plants, chromosomes): S. chaucha and S. juzepczukii. There
though a substantial amount of self-fertilizing occurs as is one pentaploid cultivated species (with 60 chromo-
well. Tubers form in response to decreasing day length, somes): S. curtilobum. There are two major subspecies
although this tendency has been minimized in commer- of Solanum tuberosum: andigena, or Andean; and tubero-
cial varieties.[18] sum, or Chilean.[21] The Andean potato is adapted to the
short-day conditions prevalent in the mountainous equa-
torial and tropical regions where it originated; the Chilean
potato, however, native to the Chilo Archipelago, is
adapted to the long-day conditions prevalent in the higher
latitude region of southern Chile.[22]
The International Potato Center, based in Lima,
Peru, holds an ISO-accredited collection of potato
germplasm.[23] The international Potato Genome Se-
quencing Consortium announced in 2009 that they had
achieved a draft sequence of the potato genome.[24]
The potato genome contains 12 chromosomes and 860
million base pairs, making it a medium-sized plant
genome.[25] More than 99 percent of all current varieties
of potatoes currently grown are direct descendants of
Potato plants a subspecies that once grew in the lowlands of south-
central Chile.[26] Nonetheless, genetic testing of the wide
variety of cultivars and wild species arms that all
After owering, potato plants produce small green fruits potato subspecies derive from a single origin in the area
that resemble green cherry tomatoes, each containing of present-day southern Peru and extreme northwest-
about 300 seeds. Like all parts of the plant except the ern Bolivia (from a species in the Solanum brevicaule
tubers, the fruit contain the toxic alkaloid solanine and complex).[7][8][9]
are therefore unsuitable for consumption. All new potato Most modern potatoes grown in North America arrived
varieties are grown from seeds, also called true potato through European settlement and not independently from
seed, TPS or botanical seed to distinguish it from the South American sources, although at least one wild
seed tubers. New varieties grown from seed can be potato species, Solanum fendleri, is found as far north
propagated vegetatively by planting tubers, pieces of tu- as Texas, where it is used in breeding for resistance to a
bers cut to include at least one or two eyes, or cuttings, a nematode species that attacks cultivated potatoes. A sec-
practice used in greenhouses for the production of healthy ondary center of genetic variability of the potato is Mex-
seed tubers. Plants propagated from tubers are clones of ico, where important wild species that have been used
the parent, whereas those propagated from seed produce extensively in modern breeding are found, such as the
a range of dierent varieties. hexaploid Solanum demissum, as a source of resistance
3
New tubers may arise at the soil surface. Since exposure fer skinning damage during harvest and handling opera-
to light leads to greening of the skins and the develop- tions. Curing allows the skin to fully set and any wounds
ment of solanine, growers cover such tubers. Commer- to heal. Wound-healing prevents infection and water-loss
cial growers achieve this by piling additional soil around from the tubers during storage. Curing is normally done
the base of the plant as it grows (hilling, or in British at relatively warm temperatures 50 to 60 F (10 to 16
English earthing up). An alternative method used by C) with high humidity and good gas-exchange if at all
home gardeners and smaller-scale growers involves cov- possible.[61]
ering the growing area with organic mulches such as straw
or plastic sheets.[60]
Correct potato husbandry can be an arduous task in some
circumstances. Good ground preparation, harrowing,
plowing, and rolling are always needed, along with a lit-
tle grace from the weather and a good source of wa-
ter. Three successive plowings, with associated harrow-
ing and rolling, are desirable before planting. Eliminating
all root-weeds is desirable in potato cultivation. In gen-
eral, the potatoes themselves are grown from the eyes of
another potato and not from seed. Home gardeners often
plant a piece of potato with two or three eyes in a hill of
mounded soil. Commercial growers plant potatoes as a
row crop using seed tubers, young plants or microtubers
and may mound the entire row. Seed potato crops are
'rogued' in some countries to eliminate diseased plants or Potato plant prior to harvest.
those of a dierent variety from the seed crop.
Potatoes are sensitive to heavy frosts, which damage them
7.1 Storage
in the ground. Even cold weather makes potatoes more
susceptible to bruising and possibly later rotting, which
Storage facilities need to be carefully designed to keep the
can quickly ruin a large stored crop.
potatoes alive and slow the natural process of decomposi-
At harvest time, gardeners usually dig up potatoes with tion, which involves the breakdown of starch. It is crucial
a long-handled, three-prong grape (or graip), i.e., a that the storage area is dark, well ventilated and for long-
spading fork, or a potato hook, which is similar to the term storage maintained at temperatures near 4 C (39
graip but with tines at a 90 angle to the handle. In larger F). For short-term storage before cooking, temperatures
plots, the plow is the fastest implement for unearthing of about 7 to 10 C (45 to 50 F) are preferred.[62][63]
potatoes. Commercial harvesting is typically done with
On the other hand, temperatures below 4 C (39 F)
large potato harvesters, which scoop up the plant and sur-
convert potatoes starch into sugar, which alters their
rounding earth. This is transported up an apron chain
taste and cooking qualities and leads to higher acrylamide
consisting of steel links several feet wide, which separates
levels in the cooked product, especially in deep-fried
some of the dirt. The chain deposits into an area where
dishesthe discovery of acrylamides in starchy foods in
further separation occurs. Dierent designs use dier-
2002 has led to many international health concerns as they
ent systems at this point. The most complex designs use
are believed to be possible carcinogens and their occur-
vine choppers and shakers, along with a blower system to
rence in cooked foods is currently under study as a possi-
separate the potatoes from the plant. The result is then
ble inuence in potential health problems.[lower-alpha 1][64]
usually run past workers who continue to sort out plant
material, stones, and rotten potatoes before the potatoes Under optimum conditions possible in commercial ware-
are continuously delivered to a wagon or truck. Further houses, potatoes can be stored for up to ten to twelve
[62]
inspection and separation occurs when the potatoes are months. When stored in homes, the shelf life is usu-
unloaded from the eld vehicles and put into storage. ally only a few weeks.[63] If potatoes develop green ar-
eas or start to sprout, these areas should be trimmed be-
Immature potatoes may be sold as new potatoes and are
fore using.[63] Trimming or peeling green areas are inad-
particularly valued for taste. These are often harvested by
equate to remove copresent toxins, and such potatoes are
the home gardener or farmer by grabbling, i.e. pulling
no longer suitable as animal food.[65][66]
out the young tubers by hand while leaving the plant in
place. Commercial storage of potatoes involves several phases:
drying of surface moisture; a wound healing phase at 85%
Potatoes are usually cured after harvest to improve skin-
to 95% relative humidity and temperatures below 25 C
set. Skin-set is the process by which the skin of the
(77 F); a staged cooling phase; a holding phase; and a
potato becomes resistant to skinning damage. Potato tu-
reconditioning phase, during which the tubers are slowly
bers may be susceptible to skinning at harvest and suf-
warmed. Mechanical ventilation is used at various points
7.3 Varieties 7
7.2 Yield
Bamberg potatoes
The blue or purple potato originated in South Amer- teins to be made, but rather prevent proteins from being
ica. It has purple skin and esh, which becomes blue made via RNA interference.[90][91][92]
once cooked. It has a slight whitish scab that seems to be
present in all samples. The variety, called Cream of the
Crop, has been introduced into Ireland and has proved 9 Pests
popular.[80] To preserve the purple color best, microwav-
ing is the best way to cook it; however, it can also be baked
Main article: List of potato diseases
and steamed. The purple or blue potato tends to have a
The historically signicant Phytophthora infestans (late
more nutty avor than its relatives of other colors.[81]
A mutation in the varieties P locus causes production of
the antioxidant anthocyanin.[82]
10 Uses
Potatoes are used to brew alcoholic beverages such
as vodka, potcheen, or akvavit.
German Bauernfrhstck
Fish and chips
In Germany, Northern and Eastern Europe, especially
In the UK, potatoes form part of the traditional staple sh in Scandinavian countries, Poland, Russia, Belarus and
and chips. Roast potatoes are commonly served with a Ukraine, newly harvested, early ripening varieties are
10.1 Culinary uses 11
considered a special delicacy. Boiled whole and served 10.1.4 North America
un-peeled with dill, these new potatoes are tradition-
ally consumed with Baltic herring. Puddings made from
grated potatoes (kugel, kugelis, and potato babka) are
popular items of Ashkenazi, Lithuanian, and Belarusian
cuisine.[114] German fries and various version of Potato
salad are part of German cuisine. Bauernfrhstck (lit-
erally Farmers breakfast) is a warm German dish made
from fried potatoes, eggs, ham and vegetables.
Cepelinai
12 See also [10] Miller, N (29 January 2008). Using DNA, scientists
hunt for the roots of the modern potato. American As-
List of potato museums sociation for the Advancement of Science. Retrieved 10
September 2008.
Loy, a form of early spade used in Ireland for the
[11] Ames, M.; Spooner, D. M. (February 2008). DNA
cultivation of potatoes.
from herbarium specimens settles a controversy about
New World crops origins of the European potato. American Journal of
Botany. 95 (2): 252257. doi:10.3732/ajb.95.2.252.
Potato battery PMID 21632349.
[1] See text: acrylamides, esp introduction; acrylamide was [15] J. Simpson; E. Weiner, eds. (1989). potato, n. Oxford
accidentally discovered in foods in April 2002 by scien- English Dictionary (2nd ed.). Oxford: Clarendon Press.
tists in Sweden when they found the chemical in starchy ISBN 0-19-861186-2.
foods, such as potato chips, French fries, and bread that
had been heated (production of acrylamide in the heating [16] David Wilton, Ivan Brunetti; p94 Word myths: debunk-
process was shown to be temperature-dependent) ing linguistic urban legends; Oxford University Press US;
2004; ISBN 0-19-517284-1
[6] University of Wisconsin-Madison, Finding rewrites the [21] Chilean Tetraploid Cultivated Potato, ''Solanum tubero-
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[7] Spooner, David M.; McLean, Karen; Ramsay, Gavin; Spooner, Univewrsity of Wisconsin, published in ''Crop
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doi:10.1073/pnas.0507400102. PMC 1253605 . PMID tuberosum L.) using morphological data and AFLP mark-
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