Institution
Albert Einstein College of Medicine
Brandeis University
Brandeis University
California State University Dominguez Hills
California State University Sacramento
Case Western Reserve University
City University of New York
Cold Spring Harbor Laboratory
Cornell University
Cornell University
DePaul University
Emory University
Georgetown School of Medicine
Gerstner Sloan Kettering
Harvard University
Harvard University
Harvard University
Harvard University
Harvard University
Harvard University
Harvard University
Harvard University
Harvard University
Harvard University
Harvard University
Harvard University
Icahn School of Medicine at Mount Sinai
Indiana University
Indiana University School of Medicine
Iowa State University
Johns Hopkins School of Medicine
Johns Hopkins School of Medicine
Johns Hopkins School of Medicine
Johns Hopkins School of Medicine
Massachusetts Institute of Technology
Medical University of South Carolina
Michigan Tech
New York University
New York University Sackler Institute
NIH Oxford-Cambridge
Northwestern University
Ohio State University
Purdue University
Rockefeller University
Rutgers University
Rutgers University
Rutgers University
Rutgers University
Rutgers University
Rutgers University
Rutgers University (Newark)
Rutgers University (Newark)
Rutgers University (Newark)
San Francisco State University
Stanford University
Stowers Institute for Medical Research
Thomas Jefferson University
University of Alabama Birmingham
University of Buffalo
University of California Berkeley
University of California Berkeley
University of California Berkeley
University of California Berkeley
University of California Davis
University of California Los Angeles
University of California San Diego
University of California San Francisco
University of California San Francisco
University of California San Francisco
University of California San Francisco
University of California San Francisco
University of California San Francisco
University of California San Francisco
University of California San Francisco
University of California San Francisco
University of California Santa Cruz
University of Chicago
University of Chicago
University of Colorado Anschutz Medical Campus Cancer Biology
University of Colorado Anschutz Medical Campus Cell Biology, Stem Cells and Development
University of Colorado Anschutz Medical Campus Molecular Biology
University of Colorado at Boulder
University of Connecticut
University of Illinois at Urbana-Champaign
University of Iowa
University of Kansas
University of Kansas
University of Kansas Medical Center
University of Massachusetts Amherst
University of Massachusetts Amherst
University of Massachusetts Amherst
University of Massachusetts Medical School
University of Michigan
University of Michigan
University of Michigan
University of Minnesota
University of Minnesota
University of Minnesota
University of Minnesota
Program GRE Not Required Program Link
Biomedical Sciences X https://www.einstein.yu.edu/education/phd/prospective-students/admissions.aspx
Molecular and Cell Biology X http://www.bio.brandeis.edu/grad/mcb/mcb_phd_rev.html
Neuroscience X http://www.bio.brandeis.edu/grad/neuro/neuro_phd.html
Biology X https://www.csudh.edu/gsr/graduate-studies/graduate-admission/#
Master's of Stem Cel Biology X https://www.csus.edu/bios/graduate%20program/ma-stemcell.html
Biomedical Sciences Training Program X https://case.edu/med/BSTP/apply.html
Biology X https://www.gc.cuny.edu/Prospective-Current-Students/Prospective-Students/Admissions#!
PhD Program X https://www.cshl.edu/education/phd-program/how-to-apply/
Biochemistry, Molecular, and Cell Biology X https://bmcb.cornell.edu/
Genetics, Genomics, and Development X https://gendev.cornell.edu/
Biological Sciences (MS) X https://csh.depaul.edu/academics/biological-sciences/graduate/biological-sciences-ms/Pages/default.aspx
Biological and Biomedical Sciences (except Population Bio) X http://www.biomed.emory.edu/
Pharmacology and Physiology X https://pharmacology.georgetown.edu/phd_applying
Biomedical Sciences X https://www.sloankettering.edu/gerstner/admissions/apply
Biological and Biomedical Sciences X http://www.hms.harvard.edu/dms/bbs/
Biological Sciences in Dental Medicine X https://gsas.harvard.edu/programs-of-study/all/biological-sciences-dental-medicine
Biological Sciences in Public Health X https://www.hsph.harvard.edu/biological-sciences/
Biological Sciences in Public Health X https://www.hsph.harvard.edu/biological-sciences/
Biophysics X https://gsas.harvard.edu/programs-of-study/all/biophysics
Chemical Biology X http://chembiophd.harvard.edu/
Immunology X http://dms.hms.harvard.edu/immunology/index.html
Molecular and Cellular Biology X https://gsas.harvard.edu/programs-of-study/all/molecular-and-cellular-biology
Neuroscience X http://dms.hms.harvard.edu/neuroscience/index.html
Organismic & Evolutionary Biology X https://gsas.harvard.edu/programs-of-study/all/organismic-and-evolutionary-biology
Systems Biology X http://sysbiophd.harvard.edu/prospective-students/admissions
Virology X http://dms.hms.harvard.edu/virology/index.html
Biomedical Science, Neuroscience X http://icahn.mssm.edu/education/phd
Biology X https://www.bio.indiana.edu/
Biomedical X https://medicine.iu.edu/education/graduate-degrees/phd/admissions/
Ecology and Evolutionary Biology X https://www.eeb.iastate.edu/admissions
Biochemistry, Cellular & Molecular Biology X http://bcmb.bs.jhmi.edu/
Cellular and Molecular Medicine Program X http://cmm.jhmi.edu/index.php/application-information/
Molecular Biophysics X http://pmb.jhu.edu/
Neuroscience X http://neuroscience.jhu.edu/graduate/apply
Biology X https://biology.mit.edu/graduate/prospective-students/application-process/
Biomedical Sciences X www.musc.edu/grad
Biological Sciences X http://www.mtu.edu/gradschool/programs/degrees/biological-sciences/
Biology X https://as.nyu.edu/biology/programs/phd.html
Biomedical Sciences X https://med.nyu.edu/research/sackler-institute-graduate-biomedical-sciences/
Scholars Program X https://oxcam.gpp.nih.gov/
Life Sciences X http://www.feinberg.northwestern.edu/sites/dgp/prospective-students/phd-program/admissions.html
Biochemistry X https://osbp.osu.edu/admissions
Chemistry X https://www.chem.purdue.edu/faq/index.php#gre
Graduate Program X https://graduateapplication.rockefeller.edu/
Biochemistry (Molecular Biosciences) X https://molbiosci.rutgers.edu/
Cell and Developmental Biology (Molecular Biosciences) X https://molbiosci.rutgers.edu/
Microbiology & Molecular Genetics (Molecular Biosciences) X https://molbiosci.rutgers.edu/
Neuroscience X http://rwjms.rutgers.edu/gsbs/programs/neuroscience/index.html
Pharmacology, Cellular & Molecular (Molecular Biosciences) X https://molbiosci.rutgers.edu/
Physiology & Integrative Biology (Molecular Biosciences) X https://molbiosci.rutgers.edu/
Cell Biology, Neuroscience and Physiology (Newark) X https://njms-web.njms.rutgers.edu/gsbs/multidisciplinary/multidisciplinary_mbgc.php
Immunity and Inflammation (Newark) X https://njms-web.njms.rutgers.edu/gsbs/multidisciplinary/multidisciplinary_i3.php
Molecular Biology Genetics and Cancer (Newark) X https://njms-web.njms.rutgers.edu/gsbs/multidisciplinary/multidisciplinary_cbnp2.php
Biology (Master's) X http://biology.sfsu.edu/content/applying-sfsu-biology-graduate-programs
Biosciences Varies by program https://biosciences.stanford.edu/prospective-students/how-to-apply/
Graduate Program X http://www.stowers.org/gradschool/admissions
Graduate School of Biomedical Sciences X https://www.jefferson.edu/university/life-sciences.html
Biomedical Sciences X http://www.uab.edu/gbs/home/admissions
Microbiology and Immunology X http://medicine.buffalo.edu/departments/micro/education/phd.html
Integrative Biology (IB) X http://ib.berkeley.edu/
Microbiology X http://plantandmicrobiology.berkeley.edu/graduate-program-microbiology
Molecular and Cell Biology X https://mcb.berkeley.edu/grad/prospective-graduates/admissions-and-recruitment
Plant Biology X http://plantandmicrobiology.berkeley.edu/plant-biology-0
Biochemistry, Molecular, Cellular and Developmental Biology * http://bmcdb.ucdavis.edu/admissions/index.html
Ecology and Evoltionary Biology X https://www.eeb.ucla.edu/grad_phd.php
Biomedical Sciences X http://biomedsci.ucsd.edu/how-to-apply/index.html
Biomedical Sciences X https://bms.ucsf.edu/
Chemistry and Chemical Biology X https://ccb.ucsf.edu/admissions
Developmental and Stem Cell Biology X https://dscb.ucsf.edu/
Integrative Program in Quantitative Biology X https://ipqb.ucsf.edu/
Neuroscience X http://neurograd.ucsf.edu/about-neuroscience-graduate-program
Oral and Craniofacial Sciences X https://dentistry.ucsf.edu/programs/oral-cranio-phd
Pharmaceutical Sciences and Pharmacogenomics X https://pspg.ucsf.edu/
Rehabilitation Science X https://ptrehab.ucsf.edu/rehabilitation-science-phd
Tetrad X https://tetrad.ucsf.edu/admissions
Molecular, Cell, and Developmental Biology X https://pbse.ucsc.edu/mcd/index.html
Bioisciences Programs X https://biosciences.uchicago.edu/programs
Ecology and Evolution X https://pondside.uchicago.edu/ee/index.html
X http://www.ucdenver.edu/academics/colleges/Graduate-School/academic-programs/cancerbiology/pages/admissions.aspx
X http://www.ucdenver.edu/academics/colleges/medicalschool/programs/CSD/Pages/home.aspx
X http://www.ucdenver.edu/academics/colleges/medicalschool/programs/Molbio/Pages/home.aspx
Ecology and Evolutionary Biology X https://www.colorado.edu/ebio/
Molecular and Cell Biology X https://mcb.uconn.edu/graduate-application-aid/
Molecular and Cellular Biology X https://mcb.illinois.edu/graduate/gradprospect/
Neuroscience X https://neuroscience.grad.uiowa.edu/
Ecology and Evolutionary Biology X https://eeb.ku.edu/how-apply
Molecular Biosciences X https://molecularbiosciences.ku.edu/
Interdisciplinary Graduate Program in Biomedical Sciences X http://www.kumc.edu/igpbs.html
Molecular and Cellular Biology X http://gpls.cns.umass.edu/mcb
Neuroscience and Behavior X http://gpls.cns.umass.edu/nsb
Plant Biology X http://gpls.cns.umass.edu/pb
Biomedical Sciences X https://www.umassmed.edu/gsbs/
Biophysics X https://lsa.umich.edu/biophysics/graduates/admissions.html
Chemical Biology X http://www.chembio.umich.edu/
Program in Biomedical Science X https://medicine.umich.edu/medschool/sites/medicine.umich.edu.medschool/files/assets/PIBS_GRE_POLICY_2018.pdf
Ecology and Evolutionary Biology X https://cbs.umn.edu/academics/departments/eeb/graduate/graduate-admissions
Biochemistry, Molecular Biology & Biophysics X https://www.med.umn.edu/education-training/graduate-programs
Integrative Biology and Physiology X https://www.physiology.umn.edu/degrees-and-programs/graduate-program
Microbiology, Immunology and Cancer Biology X http://www.micab.umn.edu/
Institution
University of Minnesota
University of Minnesota
University of Missouri
University of Montana
University of Nebraska Medical Center
University of New Mexico
University of North Carolina at Chapel Hill
University of Oklahoma Health Sciences
University of Oregon
University of Pennsylvania
University of Pennsylvania
University of Pittsburgh
University of Pittsburgh
University of Pittsburgh
University of Pittsburgh
University of Pittsburgh
University of Pittsburgh
University of Pittsburgh
University of Pittsburgh
University of Pittsburgh
University of Pittsburgh
University of Pittsburgh
University of Pittsburgh
University of South Dakota School of Medicine
University of Tennessee
University of Tennessee
University of Texas Southwestern Medical School Graduate School of Biomedical Sciences
University of Vermont
University of Vermont
University of Virginia
University of Virginia
University of Virginia
University of Washington
University of Washington
University of Washington
University of Washington
University of Wisconsin-Madison
University of Wisconsin-Madison
University of Wisconsin-Madison
University of Wisconsin-Madison
University of Wisconsin-Madison
University of Wisconsin-Madison
University of Wisconsin-Madison
Vanderbilt University
Washington State University
Washington University in St. Louis
Washington University in St. Louis
Washington University in St. Louis
Washington University in St. Louis
Washington University in St. Louis
Washington University in St. Louis
Washington University in St. Louis
Washington University in St. Louis
Weill Cornell Medicine
Program GRE Not Required Program Link
Molecular, Cellular, Developmental Biology & Genetics X https://www.med.umn.edu/education-training/graduate-programs
Pharmacology X https://www.med.umn.edu/education-training/graduate-programs
Division of Biological Sciences X http://biology.missouri.edu/graduate-studies/apply/
Organismal Biology, Ecology, and Evolution X http://hs.umt.edu/dbs/grad-programs/OBE/default.php
Interdisciplinary Graduate Program in Biomedical Sciences X https://www.unmc.edu/gradstudies/programs/phd/interdisciplinary-biomedical-sciences.html
Biology X http://biology.unm.edu/graduate/graduate-admissions.shtml
Biological and Biomedical Sciences Program X http://bbsp.unc.edu/admissions/
Graduate Program in Biomedical Sciences X http://graduate.ouhsc.edu/GraduatePrograms/PhDPrograms/GraduatePrograminBiomedicalSciences.aspx
Biology X https://biology.uoregon.edu/graduate-studies/apply/
Biology X https://www.bio.upenn.edu/graduate/
Biomedical Graduate Studies X http://www.med.upenn.edu/bgs/
Biological Sciences X http://www.biology.pitt.edu/graduate/how-apply
Biomedical Informatics X https://www.dbmi.pitt.edu/content/doctoral-degree-0
Cell Biology and Molecular Physiology X http://www.gradbiomed.pitt.edu/programs/cell-biology-and-molecular-physiology
Cellular and Molecular Pathology X http://www.gradbiomed.pitt.edu/programs/cellular-molecular-pathology
Clinical and Translational Science X http://www.icre.pitt.edu/phd/index.html
Computational Biology X http://www.compbio.cmu.edu/
Integrative Systems Biology X https://www.isb.pitt.edu/
Microbiology and Immunology X http://pmi.pitt.edu/
Molecular Biophysics & Structural Biology X http://www.mbsb.pitt.edu/
Molecular Genetics & Developmental Biology X http://www.gradbiomed.pitt.edu/programs/molecular-genetics-development-biology
Molecular Pharmacology X http://www.gradbiomed.pitt.edu/programs/molecular-pharmacology
Neurobiology X http://cnup.pitt.edu/training/graduate-program/phd-overview
Basic Biomedical Sciences X https://www.usd.edu/medicine/basic-biomedical-sciences
Ecology and Evolutionary Biology X https://eeb.utk.edu/
Microbiology X https://micro.utk.edu/
X http://www.utsouthwestern.edu/education/graduate-school/application-and-admissions/
Biology X https://www.uvm.edu/cas/biology/application-finances-support
Cellular, Molecular, and Biomedical Sciences X https://www.uvm.edu/cmb
Biomedical Sciences X http://bims.virginia.edu/
Biology X http://bio.as.virginia.edu
Ecology and Evolutionary Biology X http://eeb.virginia.edu
Biology X https://www.biology.washington.edu/programs/graduate/admissions
Genome Sciences X https://www.gs.washington.edu/academics/gradprogram/index.htm
Molecular Medicine and Mechanisms of Disease X http://www.pathology.washington.edu/academics/mmtp/phd-program/
Molecular and Cellular Biology X https://depts.washington.edu/mcb/
Cell and Molecular Biology X https://cmb.wisc.edu/admissions/
Endocrinology X https://www.med.wisc.edu/education/graduate-programs/biomedical-graduate-program-consortium/application-checklist/
Integrated Program in Biochemistry X https://ipib.wisc.edu/
Microbiology X https://microbiology.wisc.edu/ps_apply.php
Neuroscience X https://ntp.neuroscience.wisc.edu/
Nutritional Sciences X https://nutrisci.wisc.edu/graduate/
Pharmacology X https://www.med.wisc.edu/education/graduate-programs/biomedical-graduate-program-consortium/application-checklist/
Interdisciplinary Graduate Program X https://medschool.vanderbilt.edu/igp/
Biological Systems Engineering X https://bsyse.wsu.edu/graduate-program/how-to-apply/
DBBS - Developmental, Regenerative & Stem Cell Biology X http://www.dbbs.wustl.edu/divprograms/devbio/Pages/default.aspx
DBBS - Evolution, Ecology & Population Biology X http://www.dbbs.wustl.edu/divprograms/eepb/Pages/default.aspx
DBBS - Human & Statistical Genetics X http://dbbs.wustl.edu/divprograms/hsg/Pages/default.aspx
DBBS - Molecular Genetics & Genomics X http://www.dbbs.wustl.edu/divprograms/genetics/Pages/default.aspx
DBBS - Molecular Microbiology & Microbial Pathogenesis X http://www.dbbs.wustl.edu/divprograms/micro/Pages/default.aspx
DBBS - Neurosciences X http://dbbs.wustl.edu/divprograms/neuro/Pages/default.aspx
DBBS - Plant & Microbial Biosciences X http://dbbs.wustl.edu/divprograms/PlantMicroBioSci/Pages/default.aspx
DDBS - Molecular Cell Biology X http://www.dbbs.wustl.edu/divprograms/cellbio/Pages/default.aspx
All biomedical programs except Pharmacology X https://gradschool.weill.cornell.edu/admissions/apply-online
* - still require the GRE but score will not be shown to admissions committee
Updated May 8, 2019
Please email Joshua Hall (jdhall@unc.edu) or tweet to @jdhallphd for additions or corrections to this list.
References
The GRE Over the Entire Range of Scores Lacks Predictive Ability for PhD Outcomes in the Biomedical Sciences bioRxiv 2018
The Limitations of the GRE in Predicting Success in Biomedical Graduate School PLOS One 2017
Predictors of Student Productivity in Biomedical Graduate School Applications PLOS One 2017
A test that fails Nature 2014
How should we be selecting our graduate students? MBoC 2013
Multi-institutional study of GRE scores as predictors of STEM PhD degree completion: GRE gets a low mark PLOS One 2018