Vous êtes sur la page 1sur 64

1.

ANATOMÍA Y FISIOLOGÍA DEL SISTEMA NERVIOSO

Arquitectura interna del cerebro

c c c  c
  c  c  c  c c 
 c   c
  c
  cc  
c 
c  c 
  c
   cc

c c
  cc  c
 c
  c

Núcleos grises del cerebro

c 
 c   c  c
  c c  
  c c  
c  c  c c c
 cc c  c c
  c c  cc c
 cc  c cc
c c c
c
  c c c  c c  c c
c 
c c  c
 c  c  c  c   c c c 
 c  c  
 !c "c
c
  c c 
 c c c #c   c $
c 
 c
 c c
 c % c  c   c
 c  c  c 
 c 
 !c c  c c

 c &c   c 
 c c 
c   c  c  c c
 
c 
c  c
  ccc
  ccc
c c
c
  c c  c  c c  c  c c c
c
  c  c
$  c c c   c & c   c $
c 
c c c  c c  
$c
c  c
c

c
c
c
El núcleo lenticular

& c c  c 


 c c  
c  c
c  c  c  c  c  c  c %%%c

 c 'c  c  c c  c 
c c
 c  c 
 c
 c c
 c c  
$c
c  c  c  c  c   c  c
c  c   c  c
 c c (  c c  c c


$c  c %%%c 
 c
c  c 
 c
 c & c  c   c  c   c
  c c c 

c  c

 c  c c
c c
 c c c  c  c %%%c 
 c c

c c  c  c  c c


  cc c
c  c  c  c c
c c c  c c
 c  c  cc cc  c c
c  
$c
c  c   c c
c  c 
 c c  c    c &c c
 c c c c
c  c c
c
c c   c  c  c   c
 c
 c 
  c  c c  c &  c   c  c  c c
c  
 c
  c  c   c  c c  c
  c  c
 c
  c c
  cc

El talamo

& c   c  c   c  c  c 


 c 
 c c c  c c

 c  #c  c   c  c c    c  c c c


 c

  c&  c
 cc 
$c
c  c
  cc cc
 c   c  #c

Ôc  c  cc c
 cc 

$c 

c cc c c
 c
 c c   c c    c  c  c 
c c    c

 c 
c  cc !cc  c     c 

 c c c
c    c  c  c
!cc  c
 cc  cc c
c c  c
   c   c

  c   c c 
$c
 

 cc
Ôc  c  cc c
 cc 

$c 

cc c
 c  c
c c c  c c  c c   c   c 
 c c c $  c
  cc cc
Ôc  c  c c c 
 c c 

$c 
c 

 c c c
  cc c
 c  
c c cc c
cc

El núcleo caudado

Yc ccc
ccc cc c& c c c c
 c
c  c c  c  c  c  c   c $
c c   c c  c  c   c  c

 c  c  c c

 c
c cc c c 
cc
c c c  
c  c c c c c
 c c  c c  c   c
c
c
   c  c  c c c
c  c 
 c  c c c
  c c c
c c c c  c c  ccc  c

& ccc c c


 c c c
 c 
 c&c  c 

  c
   c c c c ccc
)c
c c 
c c
c cc

c   c c  
$c c
  c  c c c c c  c  c
" c  c c  c  c 
 c 
 c  c  c  c   c c  c 
 c

 c  c c 
$c c c
  cc  c c c  c  c c

  c c  c   c c  c c c


c c  
c 
c  
$c
 
 c c c
  c !c c c c  c 
 c   c c c

c  c 
 c
 c c c c  
c  c c *
c & c 
 c

 c cc
$c
 c
c cc  cc  cc c

cc
c c
 c 
c  
cc 
 
cc

El antemuro

& cc c c c cc c


  ccc c
  c


c
 cc  cc c
c c
cc c  c

LA SUSTANCIA BLANCA DE LOS HEMISFERIOS

c  
c 
c c c c   cc  cc

cc
c c cc c
c
  c c c
c
c c  c
 c c
 c 'c   c c  c   c  c
  #c c c
 c  c c
c
  c c c c c 
 c
  c
  c  c
 c  c c +  c  c c   c c  c
 c
  cc c cc c
 c
  ccc
 c c c
c   c c
  c c c
  c c c c
  c
c

c
  c c  c c   c c c  
c 
c c  c c
 c  c$
c c
 c c  cc cc c c  cc c

c
  c c  c  c c  c c  c
c 
c c c

 c  c c
  ccc  c c c c& c cc c
 #c  c  c  c c   c  c  c   c c c  
$c
 
 c

c
c
c
  cc cc c c
 c cc c
 c 
 c
c  c  c c
  c c  c
c c  c  c c c

cc c  cc

LAS FORMACIONES COMISURALES

' c   cc  c 


 cc

cc  c
c c
  c
 c
c c c  c   c& c  c c c
 c
 c c$c c
  c c
 c 
c cc c c 
cc

& c
 c
 c c
 ccccc c 
c c
 c
 cc
 c  #c  c  c c  c 
c   c   c c   c  c
 c

 ccc  c


  c c  cc
c c
 c
 c c
 c  c  c c c   c c c c
 c
 cc

" c   c  c
 c
 c c 
c c  
$c
  c c
  c c $c & c   c 
c
  c  c c  
$c
 c c

 c  c   c c c c c   c  c c c  c
 c

 c 
c,c cc c c
 c c  cc c
 
  c
 c  c
 c c  c  c c  cc
c  c 
c   c   c  c  c c  c   c $
c
c c c
 c c c
 cc(  !cc c c c 
c c
 c   cc

" c  c c  c


 c  c   c c  c c c c  c  c %%%c

 c c 
c c c c  
c 
c c c c
 c
  cc c c  c c
 c 
c c

,c c c   c c


 c c  cc  c c 
  c
c  c   cc  c
c  cc
cc c c c
  c  c   c $
c c c  c c c  c c c


 
$c c
 c

&c cc c c


 c
 cc c  c c  c  cc
 c   c 
c  c
 c  c  c   c  c c  
c
 c    c  c c c  c c c c c &  c  c
  c
 c  c c 
c c c  c  c  c
   cc c
 c   cc

c
c
c

-  c

& c
  c 
 !c  c c $ c  c c   c c   c  c

c c cc  #cc


 cc c
 c  cc c

c c c  cc c ccc c  c

.
c  c   c

  c   c c  c  c  c


  c  c c c
  ccc cc cc c
  cc c c
c c $  c

  c c
  c" c c c c

 c c 
c c

 c c  c  c c
  c 
c c 
 c
  c
  c  c c   c c   c  c   c  c
  c c
 
c  c  
 c c c   
c c  c  c Yc  c c
  c  cc  c
 c
cc c   cc c/c
c
   c c  c 0c
c c 
 c c  c 1c
c & c   c
 
 c c c #cc
 c c $  c cc c   c
c
 c c $  c   c c   c
  c &c c 
c
 c  c
  c  c  c
  c  c  c c 
c
 c c c"c c c c
  c c 
c c
 c  c
  c c c c c  c $  c c c   c

  c  c   c c  c  c   c  c  c c 


c
c c " c  c c  c   c c 
c  c   c  c
 $
 c c 
cc c
  c c c  c  cc cc
 c c 
cccc
c $  cc
 c   c $  cc c
 c  c  $
 c $  c
 c 
!c  c  c  c c  c
  c c  c  c  c
  c c c c c 
c c

 
c
 
 c  c

-  c  c  c  c  c  c  c


  c  c   c  c c  
c

cc c  
c c

Ôc c  
c 
c  c  c 
 c c   c  
 c  c  c
 
c c c
  ccc
 c c  cccccc
c
c
 !c c  cc c
 c c$ c c
 c c
 c
c
 c  c  c 
c c c  c  c c   c
 
  c c
 c  c c c
c   c c c  c  c
& c  
$cc c  
c 
c c

c
 c cc cc2!c
c

c  
c  c
 c  c  c  c
  c   c c  c
 
  c
 c c
c c  c  c   c
 c c c
c  c  c c
c
  c c c 
c   c c  c
  
$c c  c  c  c
 c  c  c  c c  c 
 c

  c  c c  c c


 c  c
c  c c  #c 
 c
 c 
 c   c 
 c   c c 
 c  c ,c   c

 c c c
 c c
 cc  c c
  cc cc
 c
 c  cc  c c c  c c

c
c
  c c   c c 3c c 'c  c  c
 c c

 #c c c c
c  c
 c c
c  
 c c c
c c
c  c   c  c
c  c c  c c
 cc c
c
 c c  c
  c  cc 
 c
 c

 
#c  c
  cc"4c

Ôc c
c 
 c c c c  c  cc  c
  c c

c  c
  c c
 c  c  c  c c
  c c
cc
 c   cc c
  cc
c
c  c

   cc"4cc c
cc c  c  cc 
cc
 c c
 c 5c c
c  
 c c   c   c   c

 cc cc c  
c 
cc  c c c c
ccc  ccc  c cc  c
  cc
"4cc
Ôc c
cc cc  c
  cc"4c c

c c  c   c
   c c  c  c  
 c c  c
   c 6  c c  c c c

c c c c c  c
c cc c  c c
cc
, c  c  c c  c c  c   c  c c c 
c
c  c c c c 
c  
$c
 c  c  c
   c c  c   c   c  c  c c c c
  c c c
 c
c c c c  c c 
c c c  
c 
c
 c c c
 c
  c c
  cc
Ôc c
c  c  c  c c   c  c ) c   c
 c  c c  
 c    c c
 c c

c
 c
 cccc
  cc
c
c c
cc c c
c c Y#c  c  c c  $c c  
 c
c  c  
  c

 cc  c
  cc"4c"
 cc  c c
 c  c  c c  
c 
c c  c c
c  c
 c  c  c    c cc
c c

 
 c
   cc cc  cc

c
c
& c
  c   c  
 c c
c c   c  c
 c & c c

 c   c c 


c c c

$c c  c  c  c    c & c
  c c
 c  c
 c c 
  c
c c cc
c c   c
  c  c
  c  c  c c  c 
 c c 5 c  c 
c   c
 c c  c  c  c 
c  c  c c  c
  c

c c
 c
cc c c
c
  c

7c c c c  c c  
 c
  c
 c
 c c c c
c 
$c  cccc

7c c
cc c c
 c
 c  c

 cc'  c  c
 c
 c c
 c c  
c  c c  
c c   c 
c c  c
 
 c c  c  c 
 c c c - c
  c c 
c
 c c
 cc c cc c
  c
 cc
 c%%%cc%+!c
  c c cc c 
 cc

ºrotuberancia o puente

'c c  c  c  c c  c  


 c  c
 c c  c

  c -   c c   c   c 


 c     c c   c
  cc ccc
 c c c
  c c c
 c

  c  c & c  c 


 c c   c

c  c  c  c

c c  c
   c &c c   
c c
 c c 
 c c
 c c  c  cc
 c+c+%c+%%cc+%%%!c cc c
  c

Bulbo raquídeo o médula oblongada

' c c c   c   c c c   


c  c  c  c
 
 !c
 c c  c c  $c c  c c c c c
  c   c & c c c c  
$c 
 c  c c c
  c
  c  c c   c c  c  
c & c 
 c  c   c 
 c

 c c  
 c %8c 8c 8%c c 8%%!c  c c  c
  c c 
c
  cc c  c c c cc c  c  cc c
  c c


cc c c  c c c c
  cc  c  c c

 c
 c
 cY  c c
 c c
 cc
 cc
 c 
  c

 c 
 
  c c    c  c
 c  c

c
c

 c   c 
 c  c $ c  c    c c   c  
 c

  c ccc

Sistema límbico

„  c c c c  c   c


 c c
 c
 c
c c  
 c
 c c c 
  c  c 
 c 5 c c
  c c   c  
c  c c c 

$c c  c c c
  c c c  c 
 c & c  c  
c c  
 c


  c  
 c c   c
c c
c
  c
   c

 c c  c
 
c
 cc c c  c
  c c
$c
c ccc

c  c  c 


 c  c
  c   c  c 
 c c c
 cc c c 
c c 
c& c c
  cc c
 c c

 cc
$cc


$cc  c  c c

c
 c   c   c
 c c 
$c c


$c c
cc c
 cc c
  cc c c

ºares craneales

9c
c cc c
  c  
cc cc  cc c  c c

 c 'c   c c c  c c  c  c  
 c c c
 c c

 c c c c c  c 


c  c c  c   c c c  c c c
c  c   c    c
 c   c 
  c c  c   c

c 
$c c c c c c c   c

c c

  cc cc:c c
c  c cc c  c
  c Y  c  c  c
 c  c c   #c    c 
 c 
 c

  c   c 
 cc   c
  c

c
c
c

Vascularización

& c  c c c  
c c c  c
  c   c  c  c  c c  c

  c ;  c   c  c
 c
c c c  c  c  c
$ c

 c cccccc c
$ccc c cc c
cc
 ccccc c
$ccc c cc c
 
$c  c  c
  c  c  c  c    c c c   c c
c   c c c c c   c  c
  c 5c  c c c  c  c

  c cc  c  c



 cc<  ccc c   cc
c
 c
 
$c c c c  c c  c  c 2!c & c =1>c  c
  c

c cc c c
   ccccc ccc c

   cc
   cc

c c
   c cc  !c cc c  
c c
c
c

c
 c  cc c c ccc

 c c c
 c
 

c c c c c
  c c c  c
c c c c  c

 c c c c2!c cc
 c
  cc
  c    c
c c c c  c  c
 c  c  c c c c  c c  c c
 c   c c   c   c $ 
 !c Y  c  c
 c  c
  c  c c c   c c  c c
 c  

c c
 c   c c
 c c
  c 
c  c c
 c c cc  c c c c
 c
  c  c c c 
 
c
c
c c   c

 cc c c


   cc c c  cc
c c   c
c  c
 c$ ccc  c c 
  !c

(  c  cc

c
c
& c c c  c  c  c
c  c  c
 c   c &c  c c
 c   c c c  c c  c c
 c  c c c   c
 c " c   c c  c  c c c c 
c  c  c c  
c
c
$c  c  c  c   c c   c c c
c 
c c
 c2!c

" c
c c ccc c  c c
 c c
c
c c  c
 c % c c  c
 c c   c  c 
cc c c  cc
  c  c  #c c c   c c  c c   c
!c c c
  c  cc cc  c&  c c
  c
 c c  c2!c

c   c ? c ccc c


 cc c c   c cc

c  c 
c c c    c c  c c c  c 
c c c
 c c
c  c c c   c c @3c  c c  c    c
c
 cc c
  cccc cc c  c2!c

c c   c cc#cc

Ôc  c

  #c c:c c  c-3cc-:cc
Ôc  c 

#c c3=c c  cY3ccY=cc
Ôc  c   #c c1c c  c 3cc 1cc
Ôc  c 
 #c c1c c  c'3cc'1cc
Ôc  c

 #c cccc

c   c  c c  c    c  c c c


c c
  c
 
c c c  c cc c
c  c2!cc

c  c  c cc


c 
c c c c
 
cccc c
c  c A1c
c 2!c Yc c
c    c  c   c
 c c
  c c
c  c& c
 c c  
c ccc
c
 c
  c c  cccc  
c 
c
 ?c c
 c
c  c 
 cc
 c c  c2!c

c   c   c  c c  c 


 c 
c  c
  c c c
 c 
 c  c  c
  c 
c  c   c  c
 c Y c c
 
$c c c c  c c  c  
c 
 c c   c
  c
   c c c
 c   c& c  c
  c
c c c
  c  c  c c   c   c   c  c  c c c

 c c c  cc  c   cc cc c cc  c
c  c cc   cc c   c
  c c cc   cc )  c

 cc
 c   c c  c c
 cc

c
c
c

Sistema nervioso periférico (SNº

Definición

& c  c c 


c c
  c c c
 cc cc  c
 c 'c c  c c 
 c c   c   c c c 
c
c c c  c c
  cc
   c  c
  c
c c  c  c 
 c c c  c c c  c 
 c 2!c 5c   c  c
 
c  c  c  c 
c   c   c  c c
$  !c c
 c
 ccc ccc c $ c
 c  
  c
c  c 
  c 
  c c c c
 
c cc c c c c  c

Ganglios

 c  c   c
 cc c c
  cc   c c c  c
 
  cc c
  c  c
  cBcY !c cc
) c
 c  cc c #c c  c
     c2!c

c   c  c c c  c    c c   c c c c  c


   cc 
$c,c cc cc  c
c c cc
c  c  c  c   c c 
c  c  c c  c c c

c  c

c   c c c  c


  c c  c  c
 c c  c
   c c  
$c 
 c  c   c 'c   c  c 
  c c c

 
$c  $
c cc   cc c cc  c

Nervios craneales y espinales

c  c
  c c    c c  c
 c
  c c
c

 c c  c & c   c  c  c
 c c 
 c %  c
  c
 c  c  ccc 
c

Y  c c  c
  c c    c  c c c $c c   c c  c
 c
 c cc c  c  c'c  c ccc c c

  c
 c  c cc
cc c c cc c c
c
c
   c  c   c c c  c c  c  
  c  c c c
 c c c c   c c $c c   c 
 c c  c  c
 
c  c  c   c & c  
c c   c c  c    c c c
c c 
c  c c 3c
 c c c c c c c c  c c
 c c cccc  c c   c  cc

 c c   c cc c  cc cc c


 c c c  c  c   c  c c c  c 
  c
 c c c  c cc

c  c
c c

c 
)c c c  c  c c  c

c  c   c   c   c c  c 
  c    c
 c
 c c c  c c c c c c  c  c   c c
 c  c  c  c
 c 5 c  c 
c c c c  c
c  c c   c 

  c &  c  c c c  c


  c
 c c  c  c c  c c c  c  c c   c c c

$c c c c  cCc
 c
 c c  c c
 c  c c   c c  c c c c  c  c   
 c
 c    cc c  c c  c  c c  $
 c

 c   c   c   c c


 c c  c  c  c  c
 c  c  c
 c c 
 c  c 
  c & c
 c c
 c c 
c c   c  c  c c  c  c c c c
c 
c  c  c  c  c   c  
  c c  c

 c c )  c  c c 


c   c c c 
$c c
 c  c  c% cc c
 c  c
 c
 c c
c 

 c 
 c  c  c

  c c   c   c
c   c c  c  c c  c c  c c
c 

 c

 c c c  c , c  c c   c c c  c

c c c c

 c 
 c    cc
 c

 cc
  c c) ccc c#c c

 c c c

 c
 c c c c  c   c  c  c   c  c c c
  c

- cc c c 


c
c c c cc c
 c 
 c
  c

 c  c  c c  c  c  c  c  c  c c c c c
  c
c
c c 
 c  c c 
 c
c  c   c   c   c
c  c 
 c c
) c c  c
 c  c  c c  c
  ccc ccc9 c

&c  c  c c c   c   c c    c c  #c  c


   ccc 
c
 cc=Dcc3c
cc c  cc c
    cc
cc c cc 
c

c
c
c

Clasificación de los nervios.

c c c
 
c c c cc cc  #cc

Ôc  c   c 


#c cc
c c   c  cc
c c Bc c 
$ c  c 
c  c c c c 
c c  c

 cc
Ôc  c   c 
#c c  c c   c  c    c c c

 c  cc
Ôc  c   c 
 #c c  c c 
c c    c c
 
 cc
Ôc  c  
 c 
 #c c  c c   c c  c 
 c  c
   c 
  c
cc

5 c c cc  cc cc  c


 c
  c 
  c
 c  c c c  c  c  c c c c c  c
  c 
c c    c 
c  c c c   c  c

cc
  !c c c c c

'c   c c  


c c c  c c c   
 c c 
$c
   cc cc c  #c c c c c c
  c
c  c
 c c  c 
  c c c c 
 c   c
 c  
 c   c    c -c  c c c c  c  c  c

 c c 
 c  c c
c c c   c c c c  
c  c c c
 c 
 c & c
 c c   c 
  c  c  c c c  c
 c c cc  cc' c

" c c c c  c


 c c c c c c c  c 
 c c
   cc c-c c
 c c  ccc
 cc c
c c

c
c
c

ºlexos Nerviosos

5c cc  c c  c c  cc c c   c c


 c
  c  c   c  c   c c c
 c c 
 c
  c  c,c
c cc  c  c  c c 
c cc
cc  c 
ccc c c  c  cc
c c c   c

Ôc "  c

 #c  c c   c c c
 c  c

  c -3cc
-Ac c c c  c   c

 c  c c  c
 c 2!c " c c c c
c  c  c -1c c c c c  c  c

 c c  c   c
 c , c   c

 c c c  c  c 2!c # c

 c cc


Ôc  c
 c cc
Ôc  c  c c
 cc
Ôc  c 

  cc
Ôc  c 
ccc
Ôc  c
 c c c

 c cc

c  c   cc  c cc  c ccc ccc


c
 c c
 c  c
 c   c c  c

 c c c c
 c
 c 
    cc cc c
c c c
 c
c c c  c ccc c
 c
 cc   c& c c

 c
 c cc c c


 c cc c
 c c c c c
  c c c$c c
 c   ccc c  c& c c
  c c
 cc cc  c c
 c c c   c c
c c 

  cc c c 

 cc c$cc c
   c

& c c 


c
c  cc cc c c   c-@cc-Ac
-c  c 
 c 
  c  c c c c c
c  

c  c  c c c

c
c
c 
c 'c c  c  c   c   c c  c  c


 ccc c
 c2!cc- c
c cc c c

c

c
 c  c  c c 
c c c c  c c c $ c
   c

ºlexo braquial

 c 
 c   c c c  c    c -1c c -:c c Y3c  c  c   c
 c 'c c 
c   c c  c c
c  c  c c
c
   c

 c  c c c   c  

c 2!c " c  c 
c c c
c
 cc c  cc c

 c cc c c

& c   c  c c c    c c   c   c , c   c  c


 c

c c  c2!c#c

Ôc  c cc


Ôc  c

 c
Ôc  c cc
Ôc  c cc
Ôc  c
  cc

5  c ccc c #c cc 



 cc c

 c

ºlexo lumbosacro

& c   c  
 c  c   c  c  c  c   c c c  c
   c   c c  c 
 c ' c  c  c c 
$c   c c
  c c c   c   c  c  c 37 @c  c  c   c  c
 c

 c c
cc c
 c  c

& c  c  c c c  c #c

Ôc  c  cc


Ôc  c  cc

c
c
" c cc c  c 
 cc cc c  c #c

Ôc  c

cc
Ôc  c  c
cc
Ôc  c  cc
Ôc  c  c  cc cc
Ôc  c cc c  c

Sistema nervioso vegetativo o autónomo (SNA

& c  c  c $  c  c c 


c c c 
 c  c  c

$ccc  c   c- c  c c c c


 c c c
 c  c  c c  c c$  c   c c cc
  #c  c 
   c  !c c  c 
    c c 
  c
 !c  c   c c  c   c   c c c c c   c
   c cc  c  c c c c 
 c c  c c

 c c c  c   c  $


c &  c   c c  c  c
  c c  c 

  c   c 7 $


 c c  7  
 c
 c c  c  cc
 c
 c c  c  c $
 c
c  c  c cc  c  c  $
 c c 
 c

c
$c c  c c$  c c c c c
$cc c$ c
c
 c  c

  c     c "c  c   c c  c

  c $
c  c  c  c 
c c  c  c  c
 
c

& c  c c 


c c   c c c

 c 
c
   cc
 c c c  $c  c c c c 

c

c  
$c c  c
  c  c c c  
$c c  c c c
  c 5 c   c
 c c 
c c 
c   
c c c 

$c c  c    c
   c& c  c c 
c c c    c c cc c

cc c c   cc

  cc  c

" c c c  c  c  c  


c
 c  c 
c c
 
$c c    c  c  c     c 
 c c 

$c c c
 

$cc cc c


$c c
 ccc cc c ) c

&c c c  c c$  c


  ccc
  c cc
  c  cc  cc cc  c c$ cc
 cc c
cc c c&ccc cc
 c c c c
c c  c c c c
  c  cc c c  c  c cc
 c
c ) cccc c  c

c
c
c
Sistema nervioso simpático

 c   c    c c c  $c 


c c c c c   c
 

c c  c c c   c   c c
 c c c c
  c  c c c  c c c   c   c " c c  c c  c
 c  c   c  $
 c c
 c  c c  c   $
 c
c

c
c c $ c c  c & c 
c  c  
 c
 c c 
  c c 
c c c 
c
c c   c c

c c c " c  c  c  c 
 c   c  c  c  c   c

 c c

c

cc  c

Sistema nervioso parasimpático

& c  c c c


  c
 c c c cc  c  cc
  c 
 cc c  c  c" c c c c  c
c c
 c c 
$c
 
c c'E5c&c c $c 
c  c  c
 $
 c c  c c  c   $
 c c
 c c c c   c
 cc c c c cc c$ c

& c  c  


c  c  
  c
c   c  c    c  c

 c
c c   c c  
$c  c  c  
c c  c   c c

c
 c c $c c  cc c c

c

c

Transmisión de los impulsos en el sistema nervioso autónomo

&c c   $c c c  c  c  c  c 


c c c
 c
 c    c  c c c  c  
c  c c


c  c c c  c   c   c 
 c  c   c c  c
 
cc  c
 
c 
c

&c   c $ c


 c  c
$c c  c  $c  c    c c  c
  c  c
c 
 c &c  c $ c c  
$c
  c c
cc c
  cc  cc cc  c  ccc  c$ c

 c
c c cc  c c c c c c c c c  c
 
!cc

c
c
 c   c  $
 c c

c  c   c c  c  c   c
c c   c c  c 
 c c  
 c c
c  c
 
  c 
 c  c ) c
  c c  c c 
c c
    c&c  c  c c  c c c
  cc cc
 cccc c    ccc
 cc c
c c 
  c c  c
  c c 
 c  c c    c 5 c c c
   c c   c 
  c c  c c
  c $
c  c c
 cc  c
  c ccc ccc  c c
cc
 c


c

& c  c  c 'Ec


 c  c  c c
 c  c  c  c
  c c  c
c  c  c 
 c  )c c  c c  c

  c c  
$c &c  c  c 'Ec
 c  c 
 c  c  c

 c
 c  c


  c 
  c c $ c 
  c c
 
 c c c 
 c  c 

  c c   c    c

 c&c
  c c  c
 c c
 c  c
  c
 $ 
 c c   c

Anatomía del sistema nervioso

&  c c
 c c  c c c
  cc c  c  c&c c
 c  c  c
  c   c c c c c  c
 c
 c
 c
   c' c c c  c #c

Ôc 5 c c


 cc

îc -  cc
îc -  c
îc Y 
c c
 c
Ôc (  c  c

' c c 


c

c
c
-  c c c cc c
cc c  c c
 c
  c  c  c c  c  
c c c c

 cc c$ c

' c c$  c c c

& c  c  c $  c  c  c 


  c  c  c   c
c
 ccc c   c  $
c-  c c ccc c
c
 cc c$ cc   c  c
 c c c

c c $c c
c 

$cc   c'c
 
c#cc

Ôc ' c c 


c
Ôc ' c c 
c

'E-c5 cc  F5 cc  c c-E'c

'%'Y&(5c E&*+%.'.c -&EY*5 c 'E-!c & c  c  c


 c  c c
 
c  c
 c c 
c   c c   c  c
 c c 
 c c  c
c  c  c    c  c

 c c  c

  c  c c  c & c
  c  c c  c

  #cc

Ôc &
 c cc c  cc c #cc
îc 9  c
   cc
îc ,
 c  cc   !cc
Ôc Y 
c
 
cc
îc ( 
 c
îc "  
cc
îc G c cc
Ôc -  c
Ôc (  c  c

5c  c  c 
 c c c c  c  c   #c  c  
 c

 c c   c
   !c  c  
 c c  c   
 c   c
 c  
cc
  !c

Y  c  c  c  c   c  c  


 c $  c 
 c c
c
  !cc c c  c  c c  c c c
 c
  c  c  c    c c  c c  c  c $  c c c
c c 
 
 c c    c ,c c 
c  c  c  c c
 cc
  ccc

,c

c
c
c  c c c c  c c   c c  c & c

 c  c  c  c c $c  c


  c  c

 c 
 c
'c c c #c

,c
 #c  c c c c  c  c
 c  c
 
  c c c c c c c  c   c  c  c 
 c c

c c  c  c  c c 


c c c  c  c
  c c
  ccc
c c
c
cc cc c
  #c

Y  c cc c
  #cc c   c cc cc  c
c c
c
 c c c c  c
  c c c  c
  c &c  c

 cc c c c c c  cc"

 cc c cc


 c c c
 c  c  c  
 c " c  c c c  c c c  
$c

 c
c  c 
  c    cc c  c   c

c c c
  cc c
 cc ccc c c
  cc c
 c " c c
c  c c
 c   c
c  c  c  c
 ccc 
$c
 cc ccc cc c
  cc c  cc c
c c c cc
c c  c
 c

Ycc c$  cc c c


 c  cc 
 c 
 c  c
c c  c
c c   cc
c c c$  !cc c
 c
  c

c c c
  cc c c c  c
  c

,c   #c 
c  c
  c c   c   c " c  c c
c  c  c

 c c  c   c c c  c c  c   c 


 c
  c c   c  c
 c  c & c c c  c  c  c

 c
  c c c 
 c cc c  cc cc
 c c  c
c  c cc

5
  c

c c c 
  c  c c  c  c c
 c  c

 c c c  c c  
c c  c

 
  c
   c & c
c c c c  c c  
 c  c  c c  !c  c
 
 c   c

"c

( c  c c  c  c c


c c
c c
) c  c  c c 
c c  c c c c c 
c

  c

&c c  
$c   c  c   c  c    c c  $c  c
 c  c &c c 
  c c c c c 
c  c  
 c
c
c
 
  c c
c  c  c
  c c c 
c   c
 ccc cc c
  c

5 c c


 cc

, c  c  c  c 


 c 
c  c c  c  c   c  c


 #c

Ôc -  #c c cc c


 cc
Ôc -  cc
Ôc Y 
c c
 cc

& c   c  
c c  c  c 
 c c c c   c  c  
 c c c
c c
  cc c  c  c  c c c 
 c c
  c c c
c c  
cc c  c c c  c c

& c
 c c  c c c
 cc c
  c c   c

-  cc

-  c c  c 
 c  c 
 c c  c c  c c  c )  c

 c c c $ c c c  c c  c 
  c   c


 c c  
 c 'c   c c c  c  
 c & c
  c

 c c c 
$c
c c cc c c c
 cc  c
 
c
 c 
 c 5c  c
  c $ c  c c =>c  c  c  c

 c c
c $ 
c cc cc
 c c=D>c c  c
'ccc c  c
   c  c cc c c c
 c  c c c  c  c c c c   c   c c  c 3Dc
c

 c
 ccc c


$cc  c  c
 c

c  c :1>c  c  c
  c c c c 
c c c
  c
  c 
c c c  cc 
c c  c c c
c
c
 cc  c  cc

c 
 c c  c  
 c c  c c  c c  c c c

c c
c  c c c  c   c c 
 c   c c


c
cc
c
 c
  cc c  cc cc
) c 
 c c
 c c  c c (  c c   c

  c& c
c
 c 
cc c
 c
 cc c
cc
 c c  c

 cc'  c& c  c
  cc

 c
c c cc  c
 cc c c c  c c
 c c
c  c c c c  c
  c c
  c 
c c  $c c c
  c  
 c
 cc

& c  c


  c c  c c c 
 c   c c  c
 c
  cc
 c c 
  cc

c
c
&c
c  c c #c

c
c
  c c  
c cc c=c$c@c cc  c c c

 cc
  c 
 c ccc cc  c
 !c,  cc c
 c   c c  c c 
c
  c  c  c @Dc 
 c  c
c c 
c  c
 c &  c   c  c  c

 
  c

   c 
c c   c c  c  c
c 
  c  c
 c c

c $  c - c c c $  c c  c   c  c
  c c

 c & c  c $  c c   c  c  c   c  c c  c

  c c  c
  c c  c   c c c
 c c '  c c $  c   c c
 c c  c cc c 
cc c
 cc*  c
c
 c 7

 c c c $  c c c

 cc c $  c  c


c
c c  cc c
 cc'  c

c  
c 
 c c
 c c   c  c   c   c
 
 cc cc c
c

, c c
 c
 c  cc  c c
c c cc c  
c

c 'c c c c   c c  c 
  c
 c

& c
 c c c  cc c   #c

Ôc Y  #c & c c  c 


 c
  c c  c   c   
 c
c c c  c cc c cc c
  cc c
 c   c
   c & c c
 c c 
$c c c
 
c c 
 c  c )  c    c c  c  c )  c
   cc  c c
cc c c
c
  cY  c  c
 c    c c
  c
 c  c  c c
c
c

 c  c cc
    !c c  c
Ôc 9   #c& c   c c  c  c c  cc c c
cc c  c c
  c& c  c c  c  cc

 c   
c 
 c c c  
$c c c  c
  ccc  c

  c c  c cc   c
    c  c c  c !c & c    c
  c c 
 cc c  
$cc  c
  ccc  c
 
  c c  c c  
c &c c c
 c  c

c
  c & c    c 
c   c
 c  
c c  c
 c  c
 c c  c  c 
 c &c 
 c  c  c

 c $
c
 c  c 
 c 
 c c c 
c
c  c
  c  c
  c  
  c c
 
c   cc c   c c c $  cc
c  c c c   c  c 
 c    7   c 5 c c

 cc c
 cc c c cc  c
  c

 cc c$  c

c
c
5
cc c
  c

c c c  c
  c  c  c  c c 
 c   c
  c
  cc  
c 
c  c 
  c
   cc

c c
  cc  c
 c
  c

E
 c  c c
  c

c 
 c   c  c
  c c  
  c c  
c  c  c c c
 cc c  c c
  c c  cc c
 cc  c cc
c c c
c
  c c c  c c  c c
c 
c c  c
 c  c  c  c   c c c 
 c  c  
 !c "c
c
  c c 
 c c c #c   c $
c 
 c
 c c
 c % c  c   c
 c  c  c 
 c 
 !c c  c c

 c &c   c 
 c c 
c   c  c  c c
 
c 
c  c
  ccc
  ccc
c c
c
  c c  c  c c  c  c c c
c
  c  c
$  cc c  cc

& c  c$


c

& c c  c 


 c c  
c  c
c  c  c  c  c  c  c %%%c

 c 'c  c  c c  c 
c c
 c  c 
 c
 c c
 c c  
$c
c  c  c  c  c   c  c
c  c   c  c
 c c (  c c  c c


$c  c %%%c 
 c
c  c 
 c
 c & c  c   c  c   c
  c c c 

c  c

 c  c c
c c
 c c c  c  c %%%c 
 c c

c c  c  c  c c


  cc c
c  c  c  c c
c c c  c c
 c  c  cc cc  c c
c  
$c
c  c   c c
c  c 
 c c  c    c &c c
 c c c c
c  c c
c
c c   c  c  c   c
 c
 c 
  c  c c  c &  c   c  c  c c
c  
 c
  c  c   c  c c  c
  c  c
 c
  c c
  cc

& c   c  c   c  c  c 


 c 
 c c c  c c

 c  c #c  c   c  c c    c  c c c


 c

  c&  c
 cc 
$c
c  c
  cc cc
 c   c  c#c

Ôc  c  cc c
 cc 

$c 

c cc c c

 c cc  c c    c c c
c c
   c 
 c  
c   c c  !c c  c
     

 c  c c c    c  c   c

!c c   c 
 c c   c c c c c  c

c
c

   c   c

  c   c c 
$c
 

 cc
Ôc  c  cc c
 cc 

$c 

cc c
 c
  c c c c  c c  c c   c   c 
 c c
c $  c  cc cc
Ôc  c  c c c 
 c c 

$c 
c 

 c c
c  cc c
 c  
c c cc c
cc

& c
 c
 
 c
c cc  cc  cc c
cc

c c
 c 
c  
cc 
 
cc

& c c

& cc c c c cc c


  ccc c
  cc


c
 cc  cc c
c c
cc c  c

5c'C'Y5E-%5cG 5E-5c,&c .'c9&(%'„&*%.'c

c  
c 
c c c c   cc  cc

cc
c c cc c
c
  c c c
c
c c  c
 c c
 c 'c   c c  c   c  c
  #c c c
 c  c c
c
  c c c c c 
 c
  c
  c  c
 c  c c +  c  c c   c c  c
 c
  cc c cc c
 c
  ccc
 c c c
c   c c
  c c c
  c c c c
  c
c

c
  c c  c c   c c c  
c 
c c  c c
 c  c$
c c
 c c  cc cc c c  cc c

c
  c c  c  c c  c c  c
c 
c c c

 c  c c
  ccc  c c c c& c cc c
 #c  c  c  c c   c  c  c   c c c  
$c
 
 cc

c
  cc cc c c
 c cc c
 c 
 c
c  c  c c
  c c  c
c c  c  c c c

cc c  cc

5'c„.*(5-%.E&'c-.(%'C*5 &'c

Yc ccc
ccc cc c& c c c c
 c
c  c c  c  c  c  c   c $
c c   c c  c  c   c  c

 c  c  c c

 c
c cc c c 
cc
c c c  
c  c c c c c
 c c  c c  c   c
c
c
   c  c  c c c
c  c 
 c  c c c
  c c c
c c c c  c c  ccc  c

ccc cc 
$c
c cc

& c
 c 
 c

& c c c  c  


 c  c  c 
 c 
 c &c  c 

  c
   c c c c ccc
)c
c c 
c c
c cc

c   c c  
$c c
  c  c c c c c  c  c
" c  c c  c  c 
 c 
 c  c  c  c   c c  c 
 c

 c  c c 
$c c c
  cc  c c c  c  c c

  c c  c   c c  c c c


c c  
c 
c  
$c
 
 c c c
  c !c c c c  c 
 c   c c c

c  c 
 c
 c c c c  
c  c c *
c & c 
 c

 c  c c
$' c   c c   c  
 c c

c c
  c
c c
  c c
c c c  c   c& c  c
 c  c
 c
 c c $c c   c c
 c 
c c c c c

cc

& c
 c
 c c
 ccccc c 
c c
 c
 cc
 c  #c  c  c c  c 
c   c   c c   c  c
 c

 ccc  c


  c c  cc
c c
 c
 c c
 c  c  c c c   c c c c
 c
 cc

" c   c  c
 c
 c c 
c c  
$c
  c c
  c c $c & c   c 
c
  c  c c  
$c
 c c

 c  c   c c c c c   c  c c c  c
 c

 c 
c,c cc c c
 c c  cc c
 
  c
 c  c
 c c  c  c c  cc
c  c 
c   c   c  c  c c  c   c $
c
c c c
 c c c
 cc(  !cc c c c 
c c
 c   cc

" c  c c  c


 c  c   c c  c c c c  c  c %%%c

 c c 
c c c c  
c 
c c c c
 c
  cc c c  c c
 c 
c c

,c c c   c c


 c c  cc  c c 
  c
c  c   cc  c
c  cc
cc c c c
  c  c   c $
c c c  c c c  c c c


 
$c c
 c

&c cc c c


 c
 cc c  c c  c  cc
 c   c 
c  c
 c   c   c  c c  
c
 c   c  cc c  cc c cc&  c c  c

c
c
 c
 c c c 
cc c  c c c   cc
c
 c   cc

-  c

& c
  c 
 !c  c c $ c  c c   c c   c  c

c c cc  #cc


 cc c
 c  cc c

c c c  cc c ccc c  c

.
c  c   c

  c   c c  c  c  c


  c  c c c
  ccc cc cc c
  cc c c
c c $  c

  c c
  c" c c c c

 c c 
c c

 c c  c  c c
  c 
c c 
 c
  c
  c  c c   c c   c  c   c  c
  c c
 
c  c  
 c c c   
c c  c  c Yc  c c
  c  cc  c
 c
cc c   cc c/c
c
   c c  c 0c
c c 
 c c  c 1c
c & c   c
 
 c c c #cc
 c c $  c cc c   c
c
 c c $  c   c c   c
  c &c c 
c
 c  c
  c  c  c
  c  c  c c 
c
 c c c"c c c c
  c c 
c c
 c  c
  c c c c c  c $  c c c   c

  c  c   c c  c  c   c  c  c c 


c
c c " c  c c  c   c c 
c  c   c  c
 $
 c c 
cc c
  c c c  c  cc cc
 c c 
cccc
c $  cc
 c   c $  cc c
 c  c  $
 c $  c
 c 
!c  c  c  c c  c
  c c  c  c c
  cccc c 
cc

 
c

 
 c  c

-  c  c  c  c  c  c  c


  c  c   c  c c  
c

cc c  
c c

Ôc c  
c 
c c c
 cc  c 
 c c c
 
c  c  c
  c c c
 c  c   c c c
c c  !c  c   c c  c
 c  c $ c  c

 c c
 c c
 c c c
c cc c
 c c   c  
  c c
 c  c c c

c   c c c  c  c & c   
$c c c  
c

c c

c
 c cc ccc
Ôc c  
c  c
 c  c  c  c
  c
  c c  c  
  c
 c c
c c  c  c
  c 
 c c c c  c  c c
c

  cc c
c  cc c  
$cc c  c
 c
 c  c  c  c c  c 
 c
  c
c
c
 c c  c c
 c  c
c  c c  #c 
 c
 c 
 c   c 
 c   c c 
 c  c
,c  c
 c c c
 c c
 cc  c c

  cc cc  c
 c  cc  c c c
 c c

c
c
  c c c   c c 3c c 'c  c  c
 c c

 #c c c c
c  c
 c c
c  
 c c c
c c
c  c   c  c
c  c c  c c
 cc c
c
 c c  c
  c  cc 
 c
 c

 
#c  c
  cc"4c

Ôc c
c  
 c  c  c  c   c   c c  c

  c c 
c  c
  c c
 c  c  c  c c

  c c c c
 c    c c  c
  c
c 
c 
c  c
   c c "4c c c 
c c c
  c   c c  
c c  c c
 c 5c c
c
 
 c c  c  c  c
 cc c
c c  
c 
cc  c c c cccc
  ccc  c cc  c
  cc"4cc
Ôc c
c c c c  c
  c c "4c c

c  c  c
   c    c c  c  c  
 c c  c
   c 6  c c
 c c c
c  c  c  c  c   c 
c  c c  c
  c  c 
c c , c  c  c c  c c  c
  c  c c c 
c c  c c c
c 
c  
$c
 c  c  c    c c  c   c
  c  c  c  c c c
  c c c  c
c c
c c  c c 
c c c  
c 
c  c  c
c
 c
  c c
  cc
Ôc c
c  c  c  c c   c  c ) c
  c  c cc 
 c   cc
 c
c

c
 c  c c c c
  c c 
c 
c c

c c  c c c c Y#c  c  c c  $c c
 
 c
c  c  
  c 
 c c   c
  c
c "4c "
 c c  c  c  c  c  c c
 
c 
c c  c c
c  c  c   c
 c    c  c c
c  c

 
 c
   cc cc  c
Ôc & c
  c  c 
 cc
c c  c c
 c& c
c
 c   c c 
c c c

$c c  c  c  c
   c & c   c  c 
 c   c
 c  c  
  c
c c
 c c
c  c    c   c  c
  c  c  c c  c

 c 5 c  c 
c   c  c c  c  c  c 
c  c
 cc c
  c
Ôc Y 
c c
 cc

c
c
Ôc & c  
c  c 
 c  c  c $
c #c  
 c c

  c  c c   
c c  c  c  c & c  
 c c

 cc c #cc
Ôc 7c cc
  cc c
 c
   c   cc  cc


c c c
cc c c
c
  cc
Ôc 7c c c c c c 
 c
  c
 c
 cc c
c c 
$c  cccc
Ôc 7c c 
c c  c  c
 c
 c   c 

 c c '  c
  c c
 c c
 c c  
c c c  
cc  c

c c  c  
 c c  c  c 
 c c c - c

  c c 
c  c c 
 c c c  c c  c

  c
 cc
 c%%%cc%+!c  c c cc c 
 cc
Ôc "  
c ccc

'c c  c  c  c c  c  


 c  c
 c c  c

  c -   c c   c   c 


 c     c c   c
  cc ccc
 c c c
  c c c
 c

  c  c & c  c 


 c c   c

c  c  c  c

c c  c
   c &c c   
c c
 c c 
 c c
 c c  c  cc
 c+c+%c+%%cc+%%%!c cc c
  c

G c c c  c c

' c c c   c   c c c   


c  c  c  c
 
 !c
 c c  c c  $c c  c c c c c
  c   c & c c c c  
$c 
 c  c c c
  c
  c  c c   c c  c  
c & c 
 c  c   c 
 c

 c c  
 c %8c 8c 8%c c 8%%!c  c c  c
  c c 
c
  cc c  c c c cc c  c  cc c
  c c


cc c c  c c c c
  cc  c  c c

 c
 c
 c2!ccY  c c
 c c
 cc
 c
c  c 
  c

 c 
 
  c c    c  c
 c  c

 c   c 
 c  c $ c  c    c c   c  
 c

  c ccc

' c  
c

„  c c c c  c   c


 c c
 c
 c
c c  
 c
 c c c 
  c  c 
 c 5 c c
  c c   c  
c  c c c 

$c c  c c c
  c c c  c 
 c & c  c  
c c  
 c


  c  
 c c   c
c c
c
  c
   c

 c c  c
 
c
 cc c c  c
  c c
$c
c cc

c
c
c  c  c 
 c  c
  c   c  c 
 c c c
 cc c c 
c c 
c& c c
  cc c
 c c

 c c 


$c c


$c c  c   c  c c
 c
  c  c
 cc
$cc


$cccc c

 cc c
  cc c c

" c
  c

9c
c cc c
  c  
cc cc  cc c  c c

 c2!c'c  cc c  cc  c c 
 cc c
 cc

 c c c c c  c 


c  c c  c   c c c  c c c
c  c   c    c
 c   c 
  c c  c   c

c 
$c c c c c c c   c

c c

  cc cc:c c
c  c cc c  c
  c Y  c  c  c
 c  c c   #c    c 
 c 
 c

  c   c 
 cc   c
  cc

+
 
$c

& c  c c c  
c c c  c
  c   c  c  c  c c  c

  c ;  c   c  c
 c
c c c  c  c  c
$ c

 c cccccc c
$ccc c cc c
cc
 ccccc c
$ccc c cc c
 
$c  c  c
  c  c  c  c    c c c   c c
c   c c c c c   c  c
  c 5c  c c c  c  c

  c cc  c  c



 cc<  ccc c   cc
c
 c
 
$c c c c cc  c c& c=1>c c  c


c c c c  c
   c c c c c  c c c  c

   cc
   cc

c c
   c cc  !c cc c  
c c
c
c

c
 c  cc c c ccc

 c c c
 c
 

c c c c c
  c c c  c
c c c c  c

 c  c  c  c c c
 c
  c c
  c     c
c c c c  c  c
 c  c  c c c c  c c  c c
 c   c c   c   c $ 
 !c Y  c  c
 c  c
  c  c c c   c c  c c
 c  

c c
 c   c c
 c c
  c 
c  c c
 c c cc  c c c c
 c
  c  c c c 
 
c
c
c c   c

 cc c c


   cc c c  cc
c c   c
c  c
 c$ ccc  c c 
  !c

(  c  cc

c
c
& c c c  c  c  c
c  c  c
 c   c &c  c c
 c   c c c  c c  c c
 c  c c c   c
 c " c   c c  c  c c c c 
c  c  c c  
c
c
$c  c  c  c   c c   c c c
c 
c c
 c

" c
c c ccc c  c c
 c c
c
c c  c
 c % c c  c
 c c   c  c 
cc c c  cc
  c  c  #c c c   c c  c c   c
!c c c
  c  cc cc  c&  c c
  c
 c c  c

c   c ? c ccc c


 cc c c   c cc

c  c 
c c c    c c  c c c  c 
c c c
 c c
c  c c c   c c @3c  c c  c    c
c
 cc c
  cccc cc c  cc

c c   c cc#cc

Ôc  c

  #c c:c c  c-3cc-:c
Ôc  c 

#c c3=c c  cY3ccY=c
Ôc  c   #c c1c c  c 3cc 1c
Ôc  c 
 #c c1c c  c'3cc'1c
Ôc  c

 #c cccc

c   c  c c  c    c  c c c


c c
  c  c

c c c  c cc c
c  cc

c  c  c cc


c 
c c c c
 
cccc c
c cA1c
cYcc
c   c c  c
 c c  c
c
c   c & c
 c  c  
c  c c c 
c  c

  c c  cccc  


c 
c
 ?c c
 cc
  c 
 cc
 c c  cc

c   c   c  c c  c 


 c 
c  c
  c c c
 c 
 c  c  c
  c 
c  c   c  c
 c Y c c
 
$c c c c  c c  c  
c 
 c c   c
  c
   c c c
 c   c& c  c
  c
c c c
  c  c  c c   c   c   c  c  c c c

 c c c  cc  c   cc cc c cc  c
c  c cc   cc c   c
  c c cc   cc )  c

 cc
 c   c c  c c
 c

#FF
 H  
F 
7 77  Fcc

Ôc c
c
c
c

EL SISTEMA NERVIOSO

INTRODUCCIÓN

'c   c  c 
  c  
 c 
 c c   c   c c  c

  c  c  c  c
 c  c c

c c c c
 c  
c c c
 
 c $ c , 
 c c  c
 c
c
  c c  c 
  c c
c c c  c  c  c  c
 c
 c ccc  c  c 

cc

 c

IY  c c   c c  c  c 



  c c c 
 c c
 
 c c  c c c&  c  ccc c
c
 c
 ccc c ccc
 cIc* HJ c

ORGANIZACIÓN BÁSICA DEL SISTEMA NERVIOSO:

, c c  c c  c 


  c  c  c  c c c c  c  c
 c  c
 c  c  c  
c c  c  c
$  c& c  cc c c c  c c$  c
 c
 c  c c c c c c  c  c  c
 c c c c  c  c
 c c 
cc

Ôc ' cE c- #'c


cc
 c c 
$c%
c c

 ccc c  c  cc

Ôc ' c E c " 


#%
c   c  c  c  c  c  c
c ccc  c 
 c$  cc c c
 cc c
c
  c ' c  c 

  c   c c c  c  c
 c c

 c 
$c   c cc c 
$c  cc

Ôc ' c E c 5$  #c & c  c 


 c c c 
$c 
 c c  c

$c  c
 c c c 
 c  c c c $ c c  c c
 c  c „ c c  c 
  c c c  
$c


cc

c
c
c  
$c c  c  c  c
 c c  c  
c  c c
 

  c $
 c c c c c   c  c 
 c c 
 c c
 c  
 c c  c  c  c
 c c  c c  c c  c
 c c 
c
cc
EL SISTEMA NERVIOSO CENTRAL:
& c cc c c c  c c-  cc c
  c
cc
ENCÉFALO#& c c  c  c  c
 c - c c c c 
 c
 c " 
 !c 
  ( 
 !c c  c 
 c   c
*  
 !& c $c
  c c  c c
 cc  c c
c   c    c c   c c c c   !c c c     c c
 cc c 
$!c c $c  c c
 c cc
  cc c
c c
cc
- c
  c

$!c
Ôc Y 
 c
K  c    c c
 c  
c

" 
 c
c

Y  c
&  c
Ôc ,
 c

&  c
c

( 
 c

c c c

"  
c c
c
Lccccccccc
*  
 c
(
 c
-  c
c

c
c
Ôc ( 
 c
G c c

cc
5  c  c    c c  c  c c   c  c

 c  c

 c 'c  c  c
 c I
  Ic c 
c 
c  c
 
 c & c 
c   c  c 
 c  c  c 
 c c  c  
c

 
cc
 cc 
 c  
cc  c !c
5 c c 
 c
  !c c  c c c
c
  c
$cc c
  c c 
  !c c  c    cc c  c  
c
c
c
  c  c
   !c c
   c#c

Ôc - c
  c  c  
c  !# c  c   c c
 c
   c c  c  c c
c 
cc

Ôc - c
 c c  
c 
!#c c c c  cc c
 c
   c c  c
  c   c  c  c c  c  
c  c

c 
cc

Ôc K  c    #Y  c  c c c c  


c  c & c
 
 cc c
 c  c%
#cc

Ôc E
 c 
 c  c c c    cc cc
Ôc E
 c
 c

Y  c c
c
c cc c  c    cc c cc
c  c
   c c c c &c  
$c
c  c  
c c

c c c  c  

 c  c  c 
 c   c
 c c  
c
c

Ôc ' c  
# c   c  c
 c  c
  c , 
c c
 c  c 
 c c c
c
 - c 
 c c  c

  cc c cc

c(  #c& c cc   cc


c cc
 cc c
c Y c  c 
 c
 c c   c   c  c   c   c  c

  c 
  cc c c c  c
  c cc

c
c
ccc
c  c c
 c  c  c ,c
* c  c
  cc
 c

-cc 5
  c cc

-c cc "c

cc
9c   c   c  

c c c
c  
  c
c  c  c
& c  
 c  c  c  c  
 c c c 
c c c c c c
 
c $ c
  c & c  c
  c  c c c

  c c c c & c  
 c c  c  
  c c c

  c  c 
c  c
c  c 
  c c  c
 c
c   c   c
c   c  c

 c  c c c  c  c c  c
   c c  c  c  c $  c &c c  c c c

  c
  cc cc


$c
c c
 c
   c
Cc
c    c c c   c  c  c   c
c  c Cc
 c  c c  c  c  
c 
!c c c  c
c  c c
 c  
c !c'c  cc c cc

Ôc 5 c   c c  #c  c  c c c c c  c   c   c



 c c c  c c c c $ c &  c   c c c c c c
  c  c  c  c
  c c  c c  
$c

c  c  c  c c

c
c  c   c  c 
 c c

 c c
 c
 c  c  c   c cc cc cc
  c c c   c c c  c  c  c  c
 c   c  c 5 c  c c
 c
  c Cc c  c
 
  c c  c
 c c c c   c $!c c c !c
c c c
 c   cc

Ôc 5 c  c c   #c


c  c   c    c     !c

 c   c
c c   c  c  c  c  c c c c c
 c c c c  c  c   c c  c c  c

 c  c  c c c   c  c     c c   c c
 cc cc
c c c
 c
cc c   cc

9 cc
cc c c  c c  c
c
c

 c c 
 c c   c c c 
 c c c  c Yc   c    c c

c 
c c c 
 c   c c 
 c    c &c 
 c
   
c  c  c  c   c c  c  c  
$c c  c c
c

5 c
  c c' cE c" 
c

% 

$#cc

Y c c
 c
 c c  c  c c cc c$ c    c
c c
 ccc  c   cc cc c cc  cc
 c
 c  c  c  
c & c   c  c  c
  c  c

  c c cc  c$  c

ccccccccc

.
$c  
#cE c
  c cc  c

cc

E c
  #c ' c 3=c  c c  c c  c c c  c  c 
 c
5  c c  c  c  
 c c c   c    c  c 
 c

 c c  c   c $
c c  c . c c 
 c  c
   c  c 
 c
 c c  c
    c c c 
  c " c   c
 c  c c c 
  c  c    c c    c & c   c
 c  c   
 c    c 
 c c
 c c   c c
 

$c

Y  c   c  c  c c  c c ) c   c c  c


 c c
c c   c  c 
 c & c  c  c  c c  c
 c c c
c  c c c
 c +c  c
$c  c  c   c c  c 
 c
"
cc c

$c c  c c$  c 


c

E c #c' c@3c cc c


c  cc ccccc
c  c
 c c  c  c
c c c c  c c c   c &  c  c
 cc c   c c  c c c 
 c  c   c c 
 c
  cc  c   !c c
 c    cc  c   !cc  c
cc c  c
 c
   c c  c

& c  c c c c c c  cc c  cc c  c  c
 cc c

 #c

" ccE c* c

c
c
-
 c
!c c-3c c-:c

,   c  !c c,3c c,3=c

 c  c !c c 3c c 1c

'
c cc  !c c'3c c'1c

-$

 c c $ c c
c c c

 c

K  c $  #%


c  c  c
 c c   c 
c c c
  c  
c  c  
c "
c  c  c  c
$  c

+ c    cc   #c

Mc  c   c
$ c c  c  c c  c c @3c  c c
 cc c c c   c c  cc cc   c c   c c
 c c c c  !c ,
  c  c
  c c  c  c c c
  c cc  c c

$c

 c  c   #c c  


$c   c  c
c  c c  c

 c   c c  c c 
c c 
$ccc cc
 
  cc

$c c cc  c c   c&  c  c


  c
c c  c c 
c c 
$c c c  c
 c
 c c c cc c   c
 c c 
c   c c c  c
 c
 c   cc

$c c cc c


c
  c$c
c 
$c 
 c c
c

c

9c c cc


  c    c&c c ccc
  cc
   c 
c c
 cc
c c    c
 !cc c c
 
  c c
$c  cc  !c

'c  c   c  c 


c  c  c  c $  c  c c
  c c
 cc c c

c

 c  c    #c "c  c  c  c


 c c
 c c  $c c


c

!c c  c c   c  c 'c  c   c  c
c
c
c  c  c  c  
c  
 c c  c 
 c   
c c

c  
$c  c

c 5c   c c 
c
  c

'c  c   c  c 


c  c  c  c $  c  
 c
c c
 c  c c
 c

cc  c   c

cc

& c  c c$  #c

5  c c
 c
 c c   $c c c  
$c c  c  c
 c  
c Y  c  c  c  c 
 c  c c 
c
 c
 c c c 
$c 
 c  c 
 c  c

c c    !c & c 'E+c
 c c
c c c $ c  c c  c c c
 c

$c
c c  c
 c c  
c 9c   c  c 'E+c c  c 
 c c c
  c   c c c 
 c

  c   
 c
 

 
 c c    c -  c  c c   c 
  c    c c
   !c  c
 c  c „
 c   c c   c 
  c  c

 c c
c  c cc cc c

!c& c
 cc c
 c  c
 c    c c   c   !c c   c 
  c

 c  c )  c   c   c c c  c c c c
 
$#c

Ôc 5c c   c$  cc
Ôc (  c  cc
Ôc Y 

 c
Ôc E
 c  
cc

c c
' cE c5$  #c, c
  c
cc
& c  c c$  c c
c$
c c  c c

c 
c  
!c c c c   c c c  c 'E-c  c
c c
$c  c c $ c 
  c
 c
c c  c 'E"c  c
c c 
  c 
 c c   c  c  c c
  c9
c   c
c  c  c   ccc c'E-c  c
  cc
ccc c$ c
  c  c  cc
cc c  c c
c  c    c c  c c 
c c $ c 
  c c
c    c

c
c
& c  c  c $  c
 c c c c 
c c c c
 
c c cc c$ c
 cc
$c c 
c

 c 
c
cc
' c c$  c 
c
5
c c c   c c   c  c  c    c   $
 c c 
c
 ccc  cc c c
   cc c  c 
c'c c
 c
c
c  c   cc cc 
c c  c
5$
c  c   c  c  c
 c c    c    c
 c c  c  c c c 

$c    c c  c c c


c   c
& c c  cc

$c c  c  c   c c'E5c

cc
c c  c  c
,c c   c    c c c 
c  c $ c 
 c c
c cc c
cc  c  c cc   c
cc  c
c c$ c
 c c  c   c
c c
 
cc c  
c  cc c
9  $
c c  c  c   c    c   c c
 c c c   c  c c  c  c c c c c   c
$  c c c c    c c    !c c  c   c
    c
c  c c c  c   c 
c  c
 
c c  c  c
 
 c c c 


$c 5

!c c c  c    c 
c
& c  c  c   
c c 
  c c  c  c c c
5

cc c  c c
 
c

c
cc
' c c5$  c'%("NY%-.c

c
c
c

cc
' cE c5$  c 
c
5
cc c  cc 
$c 
$cc c c  c   c
  $
 cc  ccccc c c   c $ 
c
5$
c c

c c 
c c 
$cIIc
c
c
cc  cc c  c  c c
cc c  cc c  $c 
c
'c
 cc c  c c
 cc$c cc c  c
Y  cc c  c$  c c  c c
c
 c c  c
 c  c 
 c ,c  c   c c  c  c  c c ) c c
 cc c   c  ccc) c) c
c  c c c  c   c  
c   c  c
 
c

 c
c c  c 'E5c 
c  c c 
  c c c 
c c c
 c    c c 

c c  c    c  c   c

 
c
cc
' c c5$  c"5*5'%("NY%-.c

c
c
c

cc
&
 cc c  
$c 
cc 
#c
& c  c c$  c 
c  
$cc c$ cc 
$c
c  c c   $c c  c c$  c
cc c cc
 c


  c  c c 
c   c
c c  #c c  c c
c  c 'E+c 5 c
  c  c c $
c c #c c  c
c c c
 cc  cc  !c
ccc c$
c
c 
 c 5c  c c  c 
$c c c  $c c 
c  
c
c   c c  c c
  c  c  
c c

c  c c


$c c cc  c c


  cc
$c
cc
' cE c5$  c

 
$c &  
$c'
c &  
$c" 
c
5 c c c  c , 
$c c c  c
' c


c c c c c

c c c c c
- 
 c



$c

c


$c
&c  c 
c 
 c c
' c
+
 

$c 
c c  c  c  
c c


   c
  
$c c c  c

c
c

  cc
c
+
 

$c
5 c    c  
 c c 5c c 


$c c
  c  
c c c 


$c c   c  c


  c  c
% c c 


$c 
c " c c 


$c c
K   c


 c c
 c 

$c c  c    c

 c

 cc  c   c   c
, 
$c c c  c - 

$c c c  c


' c
 c
  !c   !c
- c c c 


$c c 5 c c c 




$c c
' c c c c  
$c c c c


$c c
  c   c

& c  c
 c c c   c c c c   c   c  c
 
  c
c c 

$c c c   c c   $c c c  c
 c c c

$cc c
  cc c
 cc
ccc
ccc- c c
 c c  c c ccc c c
  #c
ccc!c' c c
 cc
 c c
 cc  c  c
ccc !c' c c 
cc
Ôc ' c c
 cc
 c#cc
Ôc &
 cc
Ôc -  cc
Ôc G cc
Ôc "  
cc
Ôc "
 c
   cc
Ôc -  cc
Ôc ' c  c  
c $c  c c
 c c c  c

  c c  c c  c   cc c c c #c '
c c
" 
cc
ccccccc 'c  c 
 c c c  
$c  c  c  c 
 c c c

c
 cc
c
c c  c  cc c c c

 c c
 c  c  c c c c 
  c c c c c c

c
c  c
   c cc c  c  c
cccccc & c 
 c c c c  c  c 
  #c  c   
 c c
  c
  c c 
 c c
  c cc c 
 c c
  c c
cccccc & c -  c  c c c  c c  c 
 c
 c  c   c

   c c  c  c  c c  c c  


c 
c  c

 c
 c

c
c
cccccc c -c 
 c c
c   c c
c  c
 c  c
 
c  c 'c  c c  c c   c c

 
  c  c
 c 
c c
  c
cccccc c   c c c c  c c 
 c  c   c c c  c
 c
 c  cc c
  c c
c  c c

 !c
cccccc c c
 c  c
  c  
c 
c c
c  c

 cc  
c c  c    !c
cccccc & c G c  c  c c  c
$
c c c c   
c c  c

c
c c  c  c c c  c
cccccc c "  
c c c c c
c  c  c
  c  c   c  c
 
 cc c
c c   c
cccccc& c-  c c cc c c
  c  c cc c  
cc
 c  c -  c c  c   c  c  c c  
  c  c  c & c

  c  c  c


c  c  
 c  c c 
 c
  c
  c c c   
c  c c 
 c
  c  c c  c  c
 c c
 c
  c  c
cccccc & c  c  c   c  c c  c
  c  
 c  c
  c
 c
 c  c  c

c
 c c

E c  c c  


$c c 
 c c
 c  c
c   c  c
 
$ cc

c
c
-c
 c c
 ccc 
$c
 c c
 c
  cc
c
 c
 ccc c c 
 c  c  cc c&  c  c
c  c    c  c
 c  c
 c  c c  c 
 c c
c 

$c c c   c " c
 c  c $c  c c c  
$c

c c  c c  c c c   $c c c  c  c c
$c c
 c  c c  c
  cc

c  c  c   $


7c c c  c 
  c c 
  c


  c c    c "c c  c c   c  c
 c
  c 
c c 
$c
$
ccc
c ccc cc 
  
 c
cc
c


$c

" c c
 c c  c c
 c c  
cc cc  c cc
 c c
c
c
 c c
 
$c
 cc
 ccc  cc  c c
  c

c  
c c c

c
 c c c c cc cc cc
c
 c
   cc  c c  c c  c  c c
 cc c
c  
c 
c  c  c  c 
  c c  c   c
 
  c
   c  c c   c  c  c  c c  c 
  c c
 c c   $c c c  c
 c   c c c  c c  c
 c c 
c
cc
& c   c  c c c c c  c c c  c  c $ c  c  c  c
cc c   cc c  c
c
cc
Ôc c   c $c 
 
$c   c  c c  c c  
c c   $c
 c c c  
c c   c 
c  c c

c  c   c  c

  c c 
c $ c c c

 c cc
Ôc c   c 
c c  
 c c 
c
 c 
  c c
  $c  c   c  c   c   c c c 5 c c c
5

cc
cc
c
 
$cc c c c c   c cc c   c
 cc c c c c c
 c  c
cc c
 c  c  c c c 
c  c  c   c c  
c

 c
cc
& c
$c c
 c cc cc
c c c
c
c c c 
  c  c   c c   c c  c  c  c c c

$c  
 c c  

$c c c c   c


  c )c c c
c c c 
$c c c   c 
c & c   c c c  $c
  c c
 c c  c  c  c c
 c c  
$c E c
 c c  c  c  c 
$c  c
c c  c c  c
 
c    c c  c  c c  c c $ c c  $c c  c
 c

  c
c
c
cc
c   c  

c c c  c c c  c  c c c c


c c c   c 
c c  c
 c c c c 

$c c
 $c
cc c 
$ccccc c c   c
cc
'  c   c
cc
&c c  c   c
 c c
   c  c
  c  c cc
c
ccc   c   c&c c  c c
  c

 c
 
 c 
 cc) c c  c c c
 
 c
cc
c
 c
   c c  c   c  c  c c  c  c
  cc c

cc c  c  c
 c   c&  c

 c  c   c c   c c   c c c c  c c
c 
 c c  c c 
 cc 
 c
c c 
$cc c

c c
$c  c   cc c
 c  c
cc
'  cc c  c
cc
&c c   c  c 
 c  c
 c c c $c
c c c

c  c c  c  c c   c  c c c  c c  
$c
 c  
#c c 
 c
   c & c  c  c c c c c  c

  c c  c
  c  c  c G c c "  
c c "
 c

   c c  c -  c & c  c  c c c c  c  c   7
   7 7

 !cc c c


  c c
 c
cc
& c  c  c   c c c $c
c c
c c  c c c
 c  c c  c
c c  c  c
cc c c
c c
c   c 
 c c c  c c  c c c
 
    c$c c
cc
c  
$c c  c  c
 c c  
c c  c c c c

 
$c c  c  c  c c  
  c c
 c  c
 c 5  c c  c  c c c
 c    c

c )  c
  c c  c  c

c    c 
  c c   c
 c c

  c c c c
cc
&c c  c c 
c  c
  c  
  c  c 
  c c
 c  c    c c  c   c
c c  
$c 
 c  c
 c cc c 
ccc
  c
c c $c c
c c c
 !cc c  ccc ) c

cc c c  c
 c  c 
  c   c   c   c c 
 c   #c
c    c  c  c c c 
c 
 c  c c &  c   c
  c
 c c 
 c c c  c 
c  c  c  c

 cc c c 


$cc  c  c  c  c   c

 c  c ccc c


c cc" c cc  c  c
c
c

  c c c ccc  c
  c   cc c
 c c

c c c c


$c c  c   c 


 c 5 c  c  c
 c c c 
 c c c c 
 c c c c  
c  c

 c (
 c   c 

  c  c  c  c c c  


c
   c c   c
 c c
  c   c 

 c c  c c
   ccc c c

 cc
  cc  c c 

$c
  cc c  c
cc
cc c
cc
c  c
  c c c  c c
 c c  c
 c  c  c
  c
  c c  cc  c   c  c
 c c  c   c c   c
 c 
 c
c c   c   c c  c  c   c c c
c
  c
cc
5 c  c c  c c
 c c c c  c c   c c
  c c  c 
c  c  c  c
 c c c c   c

c c c  c c c c c cc  c  c c&c

$cc cc c
 cc
cc c c c c c
 c  c   c c c c  c 
c c    c 
  c c
 c c c  c c  c  c 
c  c
  c  c   c
   cc   c
c c
  cc   c
c c ccc c 
cc
c  c  c 3=c c @3c  
c c  c c  c
  c c
  c c  c  cc 
 cc c
 c
cc  c

$#c c
c8c c c cc ccc$ c  cc c
 c
c
 c  c $c c  c   c c  $c c 
$c  c  c  c    c c  c
  c  c  c  c c  c c  c
  c %%c +%%%c c +%%c
 
c ,c c  c
  c   c c  c 
  c
   c c c
 c c  c c
c c c c c c c 
 c c

 cc c 

$cc c 
$c c c   c  c c  c
   c c c   c  c  c   c  c  
c c  c  c & c


 c   c  c   #c  c  c c c c c  c
 c   cc c  cc
c  c  c
cc
' c c c c' c c$  c
cc
& c cc  c   cc c c c cc c$ c
c c
cc  c
 c c
 c
 c c $ cc c  c

 !c&  c  c
 c c  c c cc ccc
 c 

  c
c c 
$c  c c  c $
c c
c  c  c c
c  c c  c  c  c
 c  c   c  c  c  c
 c 
c c cc c$ccc c  c  cc c

c  c 


cccc c c   c c c c
  cc c c c   c
 c

c
c
 c  c c  c c c 
c c c c
cc
 c   c c  c 
 c  c 
c c  c
  c c c c c  c c c
  c   c &  c  c 

  c
 c  c 
  c c c   c
   c

   c  cc  c
cc

-  c c c  c 


  c    c  c  c  c &c  c  c

$c
 c  c
 c  cc   c& c  c c
 c
c 

$c c  c    c  c 


  c c c   c    c

   c
  c c   c c c 
 c   c c 
 c   c c c c  c
-   c  c c
 c  c c c c  c  c
  c c   c c

 c 

c c  c 

  c  c  c
  c c
 c  c  c  c
   ccc cc
c

 c
 cc c 

$c   c
& c  c c c c
 cc c   c$
 #c
Ôc & c 
c  c  
 !c   c  c
$c  c c   c

c  c cc c c c  c c c   c
c c
c 
cc
cc
Ôc & c  
c  c
 
 !c c c 
 c   c c c  c
  cc c 
$c c $cc c cc
cc
& c 
c
  c c c
c c   c  c c   !c


 cc
c  cc c
c  ccc  c  cc
c
c
 c  c c c
 c c  c
 
c&  cc ccc c
c cc ccc c$ c c
cc

c  cc  c
 c 
 c

c
c c  c c
 cc c
c  c 

  c c c
c   c
c c   c   c  c   c  c
 
c  c c
  cc c
c c c
  cc 
 c c c
  cc c cc c  cc  ccc !c  c cc c
$ c c c  c  c
 c  c c  c   c c c  c  c
 c

c Neuroanatomiac
c

GC G.c *5OC%,&.c

c
c

c
c
c
-&*&G& .c

c
c

c
c
c
-&*&G*.c

c
c

c
c
c
&E-&„5 .K*5(5c

ccc

c
c
c
(&,C 5c &'"%E5 c

c
c

c
c
c
.;.c

c
c

c
c
c
#FFHHH  
F  P c

c
c
c
c
 c  c  c 'E-c c c c

  c c 
  c c   c c
  c c   c  c  c

  c c


  c c c c c

 c c  c
c
c
&
 cc
c
c

 
$c 
 
c c c  c   #c  c
  c c  c
    c
c
c
$c c   c c  c c c
$c c c c

 cc c
 cc c
 c cc c c c)
c c c

 c

  c c  c c  


  c   #c
7 c  c  c c
c 
c
7 c c c  c c &  c
7 c c c  ccK c c  c cc c$ cc !c
c
5 cc c  c c c c c  c
  c 7 
!cc
c  c  c  #c
 

 c
"  c c c c  c
 c c  c  c  
c  c )
c
  c  c  
  c c c  c  c   c  c c c
  c c c c  c
 c cc) c c  cc
  c  c     c  c  c  c
 c c c c c c
 
c c c c  c c  c $  c
  c  c c c c c c


c  cc c
 c cc ccc  c c  #c
 c
   c   c c  c
c
2E #c   c c c c  c   c
  c c c
  c c c

  c
 cc

$c
cc c
$ccc c cc c

 c  c c  c c c c c  c


  c c   c
c
c
c c   c   c c  c c c 
c 
c  c c
 c 0c  c   c c c
c

 c c c  c
 c  c  c
 c  c
&c c
 c
 c c  cc  c cc c
  cc
c 
!c
&c   c  c c  c  c c c  c
 c   c c c c

c
c
 
$c  c  c  c c c c c
     c  c
"%-5c c  c   cc  !ccccc c 
$cc c  c c
, Q c  c  c    c c 
c 
c c ?c c c  c c
 c  c 
c    
 c c  c  c  
c   c
c
& c 
c   c
c c c
c cc c  
c ccc
c c  c R
 c c  c  c c
 c R
c
&c  c c  c  c c c c
c  c c c

    c 5%-5!c c c c c c  c


  c  c
ccc c+%cc
 ccS c
ccc c c+%%cc+%%%cc
c  c S  
  c
&c c c  c cc c 
c   cc ?cc cc
  c
 c c c  Q c c c c  c
   c    c
&  c cT  c c cccc c c
  c%%%cc%+cc
c

, Q ccc Tcc ccc c c   c


R cc  7
  !c  c
S c  c #c
c
c
  c  77777c  c c c
Rc c
c
  c 77777c  c  c

Rc 
!c c c
Rc
c
c
  c  77777c c c 
c   cc
c

& c  ?  c c <  cc  c c     c


  c c  c  c
   c 
  c
c
& c  ?   c  c c  c $
c c  c

c c$  c ccc  c c cc


c  c ccc ccc c c   c
c c  c  ?  c c  c c c c c
    c  c  c  c  c  c
   c
  c c    c c  c
 c  c  cc
 c

 #c
7 c  c c c c  c  c
   c   c
7 c  cc c cc c
Rcccc

c  c
   c    c
c
& c  ?  c c <  c c c 
 c  c
$
 c c  c  c  c c  c
  c
c
c
 c

c
"   cc<  c
cc !c

 c  c
  c c c  c
   c c   c c c   c

  c  c @ c ?
 c c c c ?
 c   c
c
 c  c c c c c
Rc c c c c
  c c
c
 c  c c cc cc
  c  cc

-
 
 c 
  c c
c c
  #c
c
c
cc
  c  c
c c ?cc c c c cR
cc

cc c 
c Q
c c Scc
c c c  c   c c
c

c , Q c cc
cc c  c c
 c
 c c
ccc c  c c c c
  cCcc c c cc

cc9 cc c 
cc c 
 c  c  cc

  c  c c c  c c c c


c c  c   c
  c
c
cc
  cc c cc c
 cc'  cc c
c c
?  cc* c c
cc c
Rcc
 c c  cc c
cc
 c  c
  ' c c  c c ?  c c  c
 c     c c c c c c  c  c c  c  c Cc c
 c c c c c c c cccc c
c
c

cc cc  cIcc c c
  Ic ccc
c
c
-
c
c
c c
  c   c c 
c c c 
Rc  c R  c

 c c  c
S ? 
c  
c  c  c Sc   c c  c c  c  c    c c  c  c

 c
 cc c Rcc

&c
 c  c
  c c  c  c 
S 
c  c  c c  ?c c c

c  
Rc  c  
 c  
  c c c  c  c #c
7 c
  7777
 c 
Rc c c  c  c
   c
7 
Q c c 7777
 c  c c  c  c

 c
7  c ? 7777c c c  Sc c  c c "%-5c
c
&c c   c  c   c
 c  c c
c c c   ?c  c
 c 
Sc c   c c
c c c c  c c 
Rc  c
 c c c
 ccc  c    c c 
c
c
 c c
c  cc ?  cc c c ? cc

(  c   c
c
9  c 
 c c c c   c  c !c c  c  c c Rc c
c 
 c c
c c   & c c 
c  c  c 
c  c
 c  c  c  c  c
 c  c    c    c =!c 
c c c
c "%-5c
&  c  c  c   c  c   c  c  c  c    c c  c
 c    c

 c  c

 c 

  c $
 c
  cc 
 c   c  c c   c c 
c c c c
c

$cc

5 c c'E-c


cc

c
c
c
c

cc
c
c
"   cc<  cccc
c

& c  ?  c c<  cc  c c     c   c c  c  c


   c

  c
c
& c ?   c  c c c$
cc c
c c$  c cc
c
c
c  c c cc c  c ccc ccc c c   cc
c  c  ?  c c  c c c c c     c  c  c  c  c
 c
   c   c c    c c  c
 c  c  c  c


 #c
7 c  c c c c  c  c
   c   c
7 c   c c c c c c
Rc c c c 
c  c
   c
   c
c
& c  ?  c c <  c c c 
 c  c $
 c c  c  c
 cc c
  c c

#FFHHH  
FH  c

-  c
c
7c- c-  c
c 7cE
 c-  c"  c
c 7c-   cc cE
 c-  c"  c
c 7c-   c-   c
c 7c-  
  c„
   c
c 7c    c
   cc
c 7c    c
   cc
7„
  c c

$cc
c
c
& c-&*&G& .c

& c
  ccc c  c
  
c$ cc c  c
 
c
 c
 c
 cc c c  c   c$ 
c

c
c
  c  c  c  c   c  c c   c c  c c c
 cc  cc 
$c   c-
c 
cc c
  cc $  cc
  c c 
c
  c #cc

3!c& c 
c c

=!c& c  c  c

c
c
c
@!c& c   c

A!c& c c

1!c& c   cc

&  c 
c c c  cc  c  c     c c
  c

&  $
c c 
  c  c
  c c  c c  c  c & c

  c  c  c  c $ c c  c 
  c c  c
  c

  c  c   
c c c  c c Mc  c c  
$c
c  c
 c    c & c  
  c  c 
c  c $  c  c  c
  !c c

c  c c 
c c& c $c c
  c c
c 
 c

c  c   c 
 c c  
$c  c c c   
c  c 
c  c

  c  c  c 
  c & c c  c
  c c 
c c c 
c
 cc   c

- c-  c

c
c c
 c #c

3!c c
c 
 c

=!c c
cc
  cc"4c

@!c c
c c

-c 
 cc
c
c
c

- c  c  c c   c  


  c 
 c c  c c   c
  cc
c  c cc cc  c  c  c
  cc c c
c c
c 
 c c  c
  cc
 cc
  c   c cc

c  cc  c
  c   c c  
c
  c 
 c
c  c
 cc  c
  cc"4c  c
  cc
  c c  cc c
c
c c
 cc  c
  cc "4c Cc c
 cc
  cc   
c
 
  c 
 c
c 3Dc
  c c "4c  c c  c   c    c c c
cc

-cc
  cc"4c

c   c c  c
  c c "4c  c c c c 
c c c
c

  c  c
  c c  c
  c c "4c   
c

 c 
c

c  c
  ccK c c%%cc c
c c

-c cc

-c
 c cc cc
 c c

c c
 cccc c
I   Ic c  c  c   c
  c c I   Ic c  
 c
 c   c c  c
c
  c   c c   c c  c

  c  c  
c
  c 
 c
c  c 
  c   c
c  c
  cc"4c

 c
  c c K c  c %%c c c c  c c   c  c
 c c c
c

  c c  cc  c


  ccc cI   Ic

Y  $ccc c
c
  c

c
c
c
 c   c  c  c
  c

c  c 
 c c c
c

  c &  c  c c  c


  c c   c c  c c c 
 c

  c  cc cc


c c
 c 
  c

  c

  c   
  c c  
  c  c
 c  c


 c  c ) c , c c c
c
  c   c   c c c
cc cccc  c   c c  c

 c  c   c c 


ccc c  
c 
c

& c    c  c c  


c 
c
 c
c c c I  c

  Ic c c
c  c Cc    c
  c  c c  
c
  c c #cc

3!c c ccc c  c

=!c  c
  c
 cc
 c
  c  c

@!c  c  c$


 cc  c
  ccK c

A!c  c c   cc  c cc c


 ccK c

&c c   c c   cc c


 c cc c c  c c
 c
 cc c$c
 c 7$ccK c c  c

 c  c  ccc  c cc  c


  cc"4c

-c c ccc c


$c
 c c c  c cc
c
 c cc  c
  cc"4c- c c c c
c c
 c
c"4c  c c
c c  
$ccc c c c c
cc
c
c
c
 cc"4c  c  c  c
  c  c
 c  c  c
   !c
c c cc  c
  cc"4c  cc cc  cc
c c

c   c  
  c  
  c c   
  c c

 c   c    c 'c
c c c  c c c  c   c   c c
  cc  c ccc c cc c
  c
c c c
c

 c  c  c c cc c


  c c cc c
 c

c
 c ccc cc  c   c
c  c c c
 c  c
 c'E-!c  
c   c 
c  c
  c   c  c
 c
  c

c
c