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Chapter 32
Obtaining Energy
All organisms require energy to maintain their complex structure. The ultimate source of energy is the sun.
Animals are heterotrophs, depending on other organisms for food. Animals fall into one of three dietary categories:
Herbivores eat mainly autotrophs (plants and algae). Carnivores eat other animals. Omnivores regularly consume animals as well as plants or algal matter. Saprophagous animals feed on decaying organic matter.
Why We Eat
Regardless of what an animal eats, an adequate diet must satisfy three nutritional needs:
Fuel for all cellular work. The organic raw materials for biosynthesis. Essential nutrients, substances such as vitamins that the animal cannot make for itself.
Feeding Mechanisms
Exceptions include parasites that absorb nutrients that have been digested by the host:
Blood parasites Protozoan parasites Tapeworms Acanthocephalans
Suspension feeders use ciliated surfaces to create a current that draws drifting food particles into their mouths.
Filter feeding is a form of suspension feeding that involves straining food from the water as it passes through a filtering device.
Deposit feeders consume the organic matter (detritus) that accumulates on the substratum.
Many annelids simply eat the substrate, digesting organic matter. Others use appendages to gather organic deposits and move them to the mouth.
Feeding Mechanisms
Predators have evolved a variety of ways to capture, hold, and swallow prey.
Many swallow food items whole. Some have specialized teeth, beaks, or tooth-like structures. Some have highly elastic jaws and distensible stomachs to accommodate large meals. Insects have 3 pairs of appendages specialized for feeding.
Feeding Mechanisms
Only mammals can actually chew their food. Mammals have teeth that are specialized for different functions.
Incisors biting, cutting, stripping leaves. Canines seizing, piercing, tearing. Premolars & molars grinding and crushing.
Feeding Mechanisms
Herbivorous animals have evolved special devices for crushing and cutting plant material.
Snails have a radula for scraping algae or plant material. Insects have grinding & cutting mandibles. Mammals have wide corrugated molars for grinding.
Feeding Mechanisms
Fluid feeders may bite and rasp at host tissues, suck blood, and feed on contents of a hosts intestines.
Absorption is the uptake of nutrients by body cells. Elimination occurs as undigested material passes out of the digestive system.
Intracellular Digestion
In intracellular digestion, food particles are engulfed by endocytosis and digested within food vacuoles.
Protozoa, sponges.
Extracellular Digestion
Extracellular Digestion
Radiates, flatworms, & ribbon worms practice both intracellular and extracellular digestion. Extracellular digestion became emphasized with the appearance of a complete digestive tract. Digestion is almost entirely extracellular in arthropods and vertebrates.
Digestive Systems
Animals with simple body plans have a gastrovascular cavity that functions in both digestion and distribution of nutrients.
Digestive Systems
Animals with a more complex body plan have a digestive tube with two openings, a mouth and an anus. This digestive tube is called a complete digestive tract or an alimentary canal.
Digestive Systems
The digestive tube can be organized into specialized regions that carry out digestion and nutrient absorption in a stepwise fashion.
The mammalian digestive system consists of the alimentary canal and various accessory glands that secrete digestive juices through ducts.
Digestive Enzymes
Enzymes are essential in the breakdown of food into small, absorbable units.
Digestive Enzymes
Proteins are broken down into individual amino acids. Complex carbohydrates are broken down into simple sugars. Fats are reduced to glycerol, fatty acids, and monoglycerides.
Food moves through the alimentary canal by cilia, specialized musculature, or both.
The gut is lined with opposing layers of smooth muscle: a circular layer and a longitudinal layer.
Segmentation involves alternate constriction of rings of smooth muscle that move the contents around, mixing with enzymes. Peristalsis involves waves of contraction behind the food mass that move it through the gut.
Reception Conduction & Storage Grinding & early digestion Terminal digestion and absorption Water absorption and concentration of solids.
Receiving Region
Mouthparts mandibles, jaws, teeth, radula, bills. Buccal cavity mouth Muscular pharynx throat Salivary glands produce lubricating secretions that may also contain toxic enzymes or salivary enzymes to begin digestion.
Receiving Region
The Stomach
The lining of the stomach is coated with mucus, which prevents the gastric juice from destroying the cells.
The Stomach
Gastric ulcers, lesions in the lining, are caused mainly by the bacterium Helicobacter pylori.
Increasing the surface area of the intestine increases the area available for absorption.
Longer intestine Villi fingerlike projections of intestinal tissue in birds and mammals Microvilli tiny processes on intestinal cells.
The first portion of the small intestine is the duodenum, where acid chyme from the stomach mixes with digestive juices from the pancreas, liver, gallbladder, and intestine itself.
Proteases, protein-digesting enzymes. Lipases for breaking up fat. Amylase for hydrolyzing starches. Nucleases which degrade RNA & DNA into nucleotides.
The liver secretes bile into the bile duct which drains into the duodenum.
Bile is stored in the gallbladder between meals. Bile salts are important for digestion of fats.
Enzymatic digestion is completed as peristalsis moves the mixture of chyme and digestive juices along the small intestine.
Absorption of Nutrients
The small intestine has a huge surface area due to the presence of villi and microvilli that are exposed to the intestinal lumen.
Absorption of Nutrients
The enormous microvillar surface is an adaptation that greatly increases the rate of nutrient absorption.
Absorption of Nutrients
The core of each villus contains a network of blood vessels and a small vessel of the lymphatic system called a lacteal.
Absorption of Nutrients
Amino acids and simple sugars pass through the epithelium of the small intestine and enter the bloodstream.
Absorption of Nutrients
Fats are emulsified by bile salts. Micelles are tiny droplets consisting of fatty acids and monoglycerides complexed with bile salts. Micelles diffuse into epithelial cells.
Resynthesized into triglycerides and pass into the lacteals of the lymphatic system.
Undernourishment occurs in animals when their diets are chronically deficient in calories.
Mice that inherit a defect in the gene for the hormone leptin become very obese.
Regulation of Digestion
Hormones help coordinate the secretion of digestive juices into the alimentary canal.
Insulin enhances transport of glucose into body cells and stimulates storage of glucose as glycogen. Results in lower blood glucose levels.
Glucagon promotes breakdown of glycogen in the liver back into glucose which is released into the blood.
Nutritional Requirements
An animal must obtain organic carbon (from glucose) and organic nitrogen (from amino acids obtained during digestion of protein) in order to build organic molecules such as carbohydrates, lipids and proteins.
Nutritional Requirements
An animals diet must also supply essential nutrients in preassembled form. An animal that is malnourished is missing one or more essential nutrients in its diet.
Nutritional Requirements
Herbivorous animals may suffer mineral deficiencies if they graze on plants in soil lacking key minerals.
Vitamins
Vitamins are organic molecules required in the diet in small amounts. To date, 13 vitamins essential to humans have been identified. Vitamins are grouped into two categories:
Fat-soluble Water-soluble
Minerals
Minerals are simple inorganic nutrients that are usually required in small amounts.
A diet that provides insufficient amounts of one or more essential amino acids causes a form of malnutrition called protein deficiency.
Most plant proteins are incomplete in amino acid makeup. Individuals who eat only plant proteins need to eat a variety to ensure that they get all the essential amino acids.