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Most pictures taken from Hope College Leaf Litter Arthropod Key online at: http://www.hope.edu/academic/biology/leaflitterarthropods/
Phylum: Nematoda
Nematode worms - common Very long, thin, elongate body Unsegmented Pointed ends (except in hookworms see bottom picture which are too small to be seen with our scopes) Similar to some thin, shortlegged centipedes, but lack legs. Similar to many vermiform (worm-like) larval stages but usually thinner, more elongate, and distinguished by pointed ends.
Phylum: Onychophora
Velvet worms - uncommon
Long, thin, soft (velvety), unsegmented body Short, stocky/stubby legs Fairly long, soft antennae Theyre not uncommon in the leaf litter, but you dont capture them in either pitfall traps or Berlese samples very often.
Snails are easily recognized by their hard shells. Fairly common in leaf-litter samples Ive never encountered slugs in either pitfall or Berlese samples, but they occur here
Phylum Arthropoda
From here on, all organisms are in the phylum Arthropoda. Characteristics of the Arthropoda: 1) Body segmented, w/ segments usually groupd in 2-3 rather distinct regions. 2) Paired, segmented appendages (between 3-400+ pairs) 3) Bilateral symmetry (in most cases) 4) Chitinous exoskeleton, which is periodically shed and renewed as he animal grows. 5) Body has more than two cell layers, tissues and organs. 6) Body cavity a true coelom. 7) Most possesses a through straight gut with an anus (in most cases). 8) Nervous system includes a brain and ganglia. 9) Possesses a respiratory system in the form of tracheae and spiracles (in most cases). 10) Possesses a open or lacunnar circulatory system with a simple heart, one or more arteries, and no veins, (in most cases). 11) Reproduction normally sexual and gonochoristic, but can be parthenogenetic. 12) Feed on everything. 13) Found everywhere.
Spiders have two distinct body segments, eight legs, no chewing mouth parts, no wings, and no pincer-like front legs. Spiders will not be classified to family or to the groups used in visual arthropod surveys, as those groups (wolf spiders, salticids, orb weavers) are uncommon in the leaf litter
Similar to spiders but look like they have one body section. Larger and longer legs than mites.
Eight fairly short legs Body not visibly segmented into thorax and abdomen; looks oval We classify two suborders of Acari: Oribatida and Ixodida (ticks). All others are classified simply as Acari
Suborder Oribatida
Hard-bodied covered with wellsclerotized plates Often appear very round (both horizontally and vertically). Play important role in litter decomposition and soil fertility
Eight legs, plus two modified front legs with large pincers Segmented body Resemble scorpions due to the pincers, but lack the tail (and stinger), and have an oval, flattened body
Oval, dorsoventrally flattened body with platelike segments Fourteen legs Can roll into a ball
Class: Diplopoda
Millipedes - common
Millipedes have long, thin multisegmented bodies with two pairs of legs per segment. Millipedes are not classified to either order or family.
Class: Chilopoda
Centipedes fairly common
Centipedes have long, thin multisegmented bodies with one pair of legs per segment. Some look long and wormlike (like nematodes), but nematodes lack legs. Centipedes are not classified to either order or family.
Class: Pauropoda
Pauropods uncommon Elongate, slender body with 11-12 segments and 9 pairs of legs Antennae with three apical branches Small head, no eyes Similar to Symphyla, Collembola, and Psocoptera, but distinguished by having 11-12 body segments, 9 pairs of legs, and unsegmented and branched antennae
Class: Symphyla
Symphylans rare Elongate, slender body with 15-22 segments and 10-12 pairs of legs Antennae slender with many segments Well-developed, distinct head, no eyes Similar to Collembola, and Psocoptera, but distinguished by having 15-22 body segments, 10-12 pairs of legs, and many-segmented antennae
Class: Hexapoda
Insects
From here on, all insects are in the Class Hexapoda. Characteristics of the Hexapoda: 1) Body with three distinct regions: head, thorax, and abdomen. 2) One pair of antennae (rarely no antennae). 3) One pair of mandibles. 4) One pair of maxillae. 5) Three pairs of legs, one on each thoracic segment (though a few insects are legless, and some larvae have additional leglike appendages). 6) No locomotor appendages on the abdomen of the adult (except in some primitive insects). 7) Legs composed of 6 segments. 8) Abdomen consisting of 11 segments.
Order: Protura
Proturans - rare
Tiny, whitish Conical head lacking both eyes and antennae Distinguished from similar Collembola & Diplura by lack of antennae
Order: Collembola
Springtails extremely common
Small, opaque to whitish, occasionally with bluishtinted eyes Variable in shape and size All have antennae, usually many-segmented and variable in length Eyes present Many have tail, some are two-pronged Often look round-headed; body often arched Distinguished from Protura and Diplura by presence of eyes, antennae with larger segments, springtail (when present)
Order: Diplura
Diplurans fairly common Small, opaque to whitish Have fairly long antennae, usually with many small segments Eyes absent Have two caudal filaments (tails) Distinguished from Protura by presence of antennae; from Collembola by absence of eyes Also somewhat similar to bristletails and silverfish (Thysanura and Microcoryphia), but have only 2 (vs. 3) pronged tails
Order: Microcoryphia
Jumping bristletails rare
Elongated, arched body covered in scales Long antennae Three-pronged tail, with middle segment longest Distinguished from silverfish (Thysanura) by more cylindrical body, arched body, large eyes
Order: Thysanura
Silverfish - uncommon
Elongated, flattened body covered in scales Long antennae Three-pronged tail, with middle segment longest Distinguished from jumping bristletails (Microcoryphia) by flattened body, small eyes
Order: Odonata
Dragonflies, Damselflies - uncommon
In the pitfall traps youll likely only find the nymphs Odonata nymphs typically have stocky bodies, fairly big heads, and three caudal filaments (=tails) Distinguished from bristletails and silverfish from stockier, unflattened body; lack of scales; three tail filaments are approximately the same length
Order: Orthoptera
Crickets, grasshoppers, katydids - Common
Crickets are very common in pitfall traps; grasshoppers and katydids are rarely encountered Two pairs of wings (sometimes reduced) Forewings narrower than the hind wings and hardened at the base. Mandibulate mouthparts, large compound eyes, Elongated, and sometimes thickened (in Gryllidae) hind legs, for jumping
Short antennae (nearly always shorter than the body). The dorsal line of the body is usually parallel or flat/level (vs. arched in many Tettigonidae and Gryllidae)
Thickened/enlarged rear femurs with spines. Antennae variable can be very long (like Tettigonidae) Often chunky-looking Sometimes have somewhat flattened bodies.
Order: Phasmatodea
Walking sticks - rare Robust body, legs half as long as the body, no longer than the body Distinguished from threadlegged bugs (Hemiptera:Reduviidae) and stilt bugs (Hemiptera:Berytidae) by thick body and legs, and segmented body. Distinguished from preying mantids (Mantodea) by lack of enlarged, praying front arms.
Order: Dermaptera
Earwigs uncommon
Order: Isoptera
Termites - uncommon
Small, soft-bodied insects, usually pale or brown. Look meal-worm like. Live in tunnels of dirt often found on trees, branches, or vines and lianas.
Order: Blattodea
Cockroaches fairly common
Flattened, oval bodies with wings usually overlapping over the body. Often have spines on legs.
Order: Hemiptera
True bugs
Summary
Paired wings, folded over body Upper part of wings usually hardened; lower portion membranous Wings are typically parallel or overlapping. True bugs, cicadas, leafhoppers, planthoppers, aphids, whiteflies Hemiptera are fairly uncommon in leaf-litter samples; Cicadidae, Cicadellidae, Fulgoroidae, and Pentatomidae are the most likely families to be found.
Thread- Legged Bug (right). Note thin body, very thin & long legs (usually longer than body), head held upright at angle
Cicadas have large, cylindrical bodies with large eyes set widely apart on the head. Have membranous wings which are usually transparent and distinctly-veined. Very short antennae Usually 2-5 cm (but as long as 15 cm)
Similar to Cicadellidae and Fulgoroidae. Best distinguished by the laterally flattened shape with a projecting, often pointy pronotum (sometimes looks like a shark fin) Small jumping insects, 12mm or less Large pronotum usually prolonged backward over abdomen
eye
eye
nose
eye
eye
Order: Thysanoptera
Thrips relatively common
Tiny (usually 1-3mm) and very slender insects with fringed wings (not always visible) Usually brown to blackish, occasionally pale.
Order: Psocoptera
Psocids, plant lice relatively common
Very small (<6mm) Prominent head with bulging, usually bluetinted or dark eyes Long slender antennae Relatively narrow necklike prothorax When present, the wings are membranous (two pairs) and usually held tent-like over the abdomen
Order: Coleoptera
Beetles- very common
Summmary
Coleoptera have four wings, with the outer two (known as elytra) typically hardened and the inner two membranous. In most families the elytra cover most or all of the body, but in some families (e.g., Staphylinidae) parts of the body and/or inner wings exposed.
What to look for when determining family: Shape (round, oval, or elongate; flattened vs. cylindrical body) Size (width) of pronotum relative to body and head Presence/absence of beaked snout Whether body and/or elytra are hardened vs. relatively soft Whether head is visible or hidden beneath pronotum Color
NOTE: Im including here mostly those families which Ive actually encountered in pitfall or Berlese samples, and have removed many slides for foliage-dwelling beetles which are commonly found in visual arthropod surveys. But this list isnt complete refer to the other insect Powerpoint and the Study of Insects book whenever you encounter something not in this slideshow.
Elongate-oval with short, broad beak and antennae not elbowed Some have slender antennae which may be longer than the body, sim. to Cerambycidae Elytra always cover entire abdomen Small to large (0.530mm, usu. <10mm)
Very similar to Tenebrionidae Differs in having lateral posterior extensions of the prosternal process that partially closes the procoxal cavities (where legs insert to body - see diagram to right, and on p. 374 of Study of Insects)
Long beaked snout extending mostly forward (at most slightly down, vs. Curculionidae which has distinctly downward-curving beak) Also known as straight-snouted weevils.
Best distinguished from any other beetle by the extremely long, filiform (threadlike) antennae, which are often as long as or longer than the beetle's body (no other beetle family has antennae longer than ~1/2 of the body length) Distinguished from Anthribidae with long antennae by lack of a beak
Bark beetles subfamily Scolytinae are abundant in virtually every pitfall sample Very blocky to me they look like little teddy bears Have short, bent, clubbed antennae and short, stout beaks, and are light to dark brown in color; usually 2-3mm long
Often called pinchingbugs because of large mandibles of the males Mandibles half as long as the body in some species Similar to Scarabaeidae, but terminal antennomeres not held as tightly together
Tiny (<1mm), oval, convex, shining, blackish body Have large, prominent head and capitate (clubbed) antennae Distinguished from Leiodidae and Phalacridae by smaller size, more distinctly capitate (slighty clubbed) antennae, dilated (flattened) tibiae
Relatively large (335mm) with soft and somewhat flattened bodies and clubbed antennae Most common species are black with yellow or orange on the pronotum
Black or brown, with filiform antennae Pronotum is nearly as wide as elytra Usually on forest floor (common in leaf litter) Many produce odorous chemicals Distinguished from Carabidae by having a pronotum which is nearly as wide as the elytra; from Scarabaeidae by filiform antennae
Elongate, medium to large (2.6-20mm); most brown to black, but some are bluish or greenish Distinguished by narrow waist, pronotum separated widely from the base of the elytra
Fairly large, with extremely hard shell, which is often bumpy Head much narrower than the pronotum Short, stout antennae Open procoxal cavity (where legs insert to body - see diagram to right, and on p. 374 of Study of Insects)
Winged forms distinguished from Diptera by presence of two pairs of wings; narrow waist; and distinct separation between thorax and abdomen. We only classify ants (family: Formicidae) to family level all other Hymenoptera are not identified to family. Family: Formicidae (ants) =>
Order: Lepidoptera
Moths, butterflies Caterpillars are relatively common, pupae uncommon, eggs/adults rare
Order: Mecoptera
Scorpionflies, Hangingflies - uncommon
Mid-sized (9-25mm), slender-bodied insects with long beak below head Four long, narrow wings with dense venation End of abdomen often with curved, bulbous appendage in males, which looks like scorpions stinger
Order: Diptera
Flies - common One pair of wings Thorax and abdomen are usually fused and approximately the same width Also have a pair of halteres, derived from the hind wings, on the metathorax (often hard to see). Most similar to Hymenoptera, but can be distinguished by the presence of only two wings, and the absence of a narrow waist separating the thorax and abdomen.
Insect Larvae
Orders: All (enter as Larvae) Common
Insect larvae are incredibly variable can look like caterpillars, maggots, worms, or grubs With a few rare exceptions (e.g., Neuroptera larvae) I dont attempt to identify order or family of larvae just record as larvae In the family column, enter the type or form of the larvae: vermiform (maggotor worm-like), scarabeiform (grub-like), elateriform (long and thin), campodeiform (grublike with bristletails), or eruciform (caterpillarlike) see diagrams on p. 47 in Study of Insects Be careful to distinguish nematodes, millipedes, and other adult insects which may appear superficially larvaelike