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Lamarck rejected fixity He proposed a theory of evolution which is attractive but it was eventually rejected because of the way inheritance works
Lamarck
LAMARK
Disproving Lamarck
Characteristics acquired during the lifetime of a parent are not passed onto the offspring An athlete who develops a large muscle mass through training does not have children who already possess this large muscle mass Ernst Haeckel In an attempt to disprove Lamarckism he is said to have cut off the tails of mice for several generations. The babies born from this line of tailless mice still grew tails as long as their ancestors. This was not exactly a fair test as the mice had not stopped using their tails in an attempt to adapt to their environment. They still found their tails useful
learned behaviour patterns can be changed within a generation Members of a social group who have acquired the behaviour in their lifetimes will pass these learned skills onto others including their children
Natural Selection
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Cactus eater
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2006-2007
Bud eater
Charles Darwin
1831-1836 (22 years old!) makes many observations of nature main mission of the Beagle was to chart South American coastline
Robert Fitzroy
Galapagos
Recently formed volcanic islands. Most of animals on the Galpagos live nowhere else in world, but they look like species living on South American mainland.
species
Darwin asked: Why were these creatures found only on the Galapagos Islands?
Alfred Wallace
Family of Modest Means Limited education, worked many jobs: construction, surveying, assistant to a watchmaker, teacher... Loved natural history and the outdoors With naturalist Henry Bates, traveled to Brazil, explored the Amazon, sold collections of biological specimens, especially insects
1848: Wallace and Bates sailed for South America, gathered large collections 1852: Sailed back to England, ship caught fire and sank after 3 weeks! Passengers rescued but collections lost! 1859: Bates gathered more than 14,000 species in Brazil 1854: Wallace collected in the Malay Archipelago & recognized the Archipelago as biologically divided by a narrow strait, separating Asian fauna from Australian fauna, still recognized as Wallace's Line.
Applied geological idea of uniformitarianism to biology Charts of a species might look more like a tree than a straight line No species came into existence unless coexisted with another similar species that was its predecessor 1855: Wrote paper arguing for evolution of species, searching for a
June 1858: Wallace drafted his thoughts, sent them in letter to Darwin, Darwin was shattered
July 1858: Joint credit was given to Darwin & Wallace at Linnaean Society The announcement attracted no particular scientific or public attention! 1859: Darwin worked intensely for the next year on his manuscript, On the Origin of Species by Means of Natural Selection (1859), sold out on day of
Natural Selection
Theory of Natural Selection was based on: Observation 1 organisms are potentially capable of producing large numbers of o spring. Observation 2 There is a competition for resources Observation 3 All living things vary. Di erent members of the same species are all slightly di erent Deduction 1 There is a universal struggle for survival. More organisms of each kind are born than can possibly obtain food and survive. Individuals with some kind of advantage have the best chance of surviving and reproducing their own kind
Observation 3 All living things vary. Dierent members of the same species are all slightly dierent
Deduction
There is a universal struggle for survival. More organisms of each kind are born than can possibly obtain food and survive. Individuals with some kind of advantage have the best chance of surviving and reproducing
Biston betularia
There are mutated individuals with dark pigmentation Formas melnicas: Carbonaria (CC) Insularia (Cc) Forma silvestre: Tpica (cc)
Around 1850, during industrial revolution, atmosphere pollution reduced the presence of lichens The trunks of trees turned dark colour
As time has passed, pollution has been reduced Lichens came back to the tree trunks and the situation changed again
The white moth is now the most common one, as it was before.
Suppose there is a population of rabbits. The color of the rabbits is governed by two incompletely dominant traits: black fur represented by B and white fur represented by b. A rabbit with the genotype of BB would have a phenotype of black fur, a genotype of Bb would have gray fur (a display of both black and white) and a genotype of bb would have a phenotype of white fur. If this population of rabbits were put into an area that had very dark black rocks as well as very white colored stone, What kind of selection would you predict?
Modes of selection
Types of evolution
1. Divergent evolution 2. Convergent evolution
Divergent Evolution
1. Divergent evolution
Two different populations of a species that experience different selection pressures may as a result become genetically different from one another
Once in a very great while, the two populations will become so different that they cannot reproduce together anymore (this then becomes speciation).
Divergent Evolution
http://www.biology-online.org/images/darwin_finches.jpg
Divergent Evolution
Divergent Evolution
Divergent Evolution
Convergent Evolution
2. Convergent evolution
Start with two very different species; they experience similar selection pressures and through time come to look and/or behave similarly.
Convergent Evolution
Fish Reptile Mammal What is the similar selection pressure?
Convergent Evolution
Convergent Evolution
What is the similar selection pressure?
http://www.votawphotography.com /photo/Animals/animals.htm
Speed to catch prey (cheetah) and speed to win races and be bred.
http://www.hundekosmos.de/images/greyhound_517.jpg
Convergent Evolution
Placental Mammals Marsupial Mammals
They observed that the fossil record gives a different picture for the evolution They claim that there were long periods of stasis (410 million years) involving little evolutionary change Then occasional rapid formation of new species As little as 5,000 - 50,000 years
Stephen J Gould
Niles Eldredge
http://anthro.palomar.edu/synthetic/images/graph_of_ph
http://anthro.palomar.edu/synthetic/images/graph_of_punctuated_equilibrium_2.gif
Homework
State what mechanisms of evolution do you know and describe it in one paragraph